Quercus coccinea |
Quercus gambelii |
|
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scarlet oak |
Gambel oak |
|
Habit | Trees, deciduous, to 30 m; lower trunk without stubs of dead branches. | Shrubs or trees, deciduous, shrubs sometimes clumped and spreading, trees small or moderately large. |
Bark | dark gray to dark brown, irregularly fissured with scaly ridges, inner bark orangish pink. |
gray or brown, scaly. |
Twigs | reddish brown, (1-)2-3.5 mm diam., glabrous. |
brown or reddish brown with few, inconspicuous lenticels, 1.5-2.5 mm diam., glabrous or stellate-pubescent. |
Buds | brown, ovoid, ca. 3 mm, apex acute or obtuse, sparsely pubescent, becoming glabrate. |
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Leaves | blade elliptic to ovate or obovate, 70-160 × 80-130 mm, base obtuse to truncate, margins with 5-9 deep lobes and 18-50 awns, lobes distally expanded, sinuses usually extending more than 1/2 distance to midrib, apex acute; surfaces abaxially glabrous except for minute axillary tufts of tomentum, adaxially glossy light green, glabrous, secondary veins raised on both surfaces. |
blade elliptic to obovate or oblong, deeply to shallowly 4-6-lobed, (40-)80-120(-160) × (25-)40-60(-100) mm, membranous, base truncate to cuneate, margins entire or coarsely toothed, lobes oblong, rounded or subacute, sinuses acute or narrowly rounded at base, reaching more than 1/2 distance to midrib, secondary veins 4-6 on each side, each passing into lobe, branched, apex broadly rounded; surfaces abaxially dull green, sometimes glaucous, densely velvety with erect 4-6-rayed hairs, sometimes glabrate or persistently villous only near midribs, secondary veins prominent, adaxially lustrous dark green, appearing glabrate, microscopically pubescent, secondary veins slightly raised. |
Acorns | biennial; cup turbinate to hemispheric, 7-13 mm high × 16.5-31.5 mm wide, covering 1/3-1/2 nut, outer surface light to dark reddish brown, glossy, glabrous to puberulent, inner surface light brown, glabrous, occasionally with ring of pubescence around scar, scales often tuberculate, base broad, glossy, margins strongly concave with tips tightly appressed, acute to attenuate; nut oblong to subglobose, 12-22 × 10-21 mm, glabrous, with 1 or more rings of fine pits at apex, scar diam. 6.5-13.5 mm. |
solitary or paired, subsessile or on peduncle to 10(-30) mm; cup deeply cup-shaped, 5-8(-17) mm deep × 7-15(-25) mm wide, enclosing 1/4-1/2 nut, base round, margin thin, scales closely appressed, ovate, markedly tuberculate, proximally gray-tomentulose; nut light brown, ovoid to ellipsoid, (8-)12-15(-33) × 7-12(-18) mm. |
Cotyledons | distinct. |
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Terminal | buds dark reddish brown, conic to ovoid, 4-7 mm, noticeably 5-angled in cross section, usually silvery- or tawny-pubescent distal to middle. |
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2n | = 24. |
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Quercus coccinea |
Quercus gambelii |
|
Phenology | Flowering spring. | Flowering mid-late spring. |
Habitat | Poor soils, well-drained uplands, dry slopes, and ridges, occasionally on poorly drained sites | Montane conifer, oak-maple, and higher margins of pinyon-juniper woodlands |
Elevation | 0-1500 m (0-4900 ft) | 1000-3030 m (3300-9900 ft) |
Distribution |
AL; AR; CT; DC; DE; GA; IL; IN; KY; MA; MD; ME; MI; MO; NC; NH; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WI; WV
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AZ; CO; NM; NV; OK; TX; UT; WY; Mexico (Chihuahua, Coahuila, and Sonora)
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Discussion | Trees having acorns with broad, distinctly warty cups are sometimes classified as Quercus coccinea var. tuberculata Sargent. Quercus coccinea reportedly hybridizes with Q. imbricaria, Q. ilicifolia (= Q. ×robbinsii Trelease), Q. laevis, and Q. palustris (E. J. Palmer 1948) and with Q. phellos, Q. rubra, and Q. velutina (= Q. ×fontana Laughlin). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Numerous hybrids of Quercus gambelii with various scrub oaks have been reported, including Q. grisea and Q. turbinella. Such hybrids in general have shallowly lobed or dentate, semipersistent leaves and intermediate characteristics of pubescence and fruit between parental types. Such hybrids are usually referred to as the Quercus ×undulata complex because of widespread application of the latter name to various populations. One population from San Juan County, Utah, with larger fruit but otherwise not differing from typical Quercus gambelii, has been recognized as Q. gambelii var. bonina Welsh. Unless other characters are found to support this segregation, the plants are best not treated as a formal taxon, particularly considering the extensive variation and hybridization associated with Q. gambelii throughout its range. Numerous putative hybrid swarms occur throughout the range of Quercus gambelii that involve a number of suspected parental species. Most of these populations have, at one time or another, been referred to Quercus undulata Torrey. The putative hybrids have serrate or shallowly lobed leaves and considerable variation in habit, leaf pubescence, and acorn morphology. J. M. Tucker (1961, 1969, 1971) and J. M. Tucker et al. (1961) have identified the major components of the Q. undulata complex as Q. turbinella (western Utah and northwestern Arizona, and central Colorado), Q. grisea (New Mexico and southern Colorado), Q. havardii (southeastern Utah and northwestern Arizona), Q. mohriana (northeastern and southern New Mexico), Q. arizonica (central Arizona), and Q. muhlenbergii (eastern and central New Mexico). Quercus macrocarpa has been implicated as a parent of variable populations in New Mexico (J. M. Tucker and J. R. Maze 1966). Because of the complex variability in these populations, no effort has been made to treat them separately here; indeed, it would be impossible to produce usable keys if these were included as formal taxa. Hybrids derived from Quercus gambelii and an evergreen species are often semideciduous, retaining a variable portion of green or brownish leaves over the winter. Quercus gambelii was used medicinally by the Navaho-Ramah to alleviate postpartum pain, as a cathartic, as a ceremonial emetic, and as a life medicine (D. E. Moerman 1986). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 3. | FNA vol. 3. |
Parent taxa | Fagaceae > Quercus > sect. Lobatae | Fagaceae > Quercus > sect. Quercus |
Sibling taxa | ||
Synonyms | Q. coccinea var. tuberculata | Q. douglasii var. gambelii, Q. gambelii var. gunnisonii, Q. lesueuri, Q. marshii, Q. novomexicana, Q. undulata var. gambelii, Q. utahensis |
Name authority | Münchhausen: Hausvater 5(1): 254. (1770) | Nuttall: J. Acad. Nat. Sci. Philadelphia 1(2): 179. (1848) |
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