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scarlet oak

canyon live oak, gold cup live oak, maul oak

Habit Trees, deciduous, to 30 m; lower trunk without stubs of dead branches. Trees or shrubs, trees small to medium-sized, to 25 m, shrubs of variable size.
Bark

dark gray to dark brown, irregularly fissured with scaly ridges, inner bark orangish pink.

Twigs

reddish brown, (1-)2-3.5 mm diam., glabrous.

branching at 60° angles or less, golden brown, 1-2 mm diam., flexible, densely pubescent 1st year, moderately so 2d year.

Leaves

blade elliptic to ovate or obovate, 70-160 × 80-130 mm, base obtuse to truncate, margins with 5-9 deep lobes and 18-50 awns, lobes distally expanded, sinuses usually extending more than 1/2 distance to midrib, apex acute;

surfaces abaxially glabrous except for minute axillary tufts of tomentum, adaxially glossy light green, glabrous, secondary veins raised on both surfaces.

blade oblong, acuminate, usually flat to slightly concave, 20-70 × 10-35 mm, thick, leathery, base obtuse to rounded, secondary veins 12 or more pairs, branching at ca. 50° angles, slightly raised abaxially, margins often slightly revolute with moderately thickened cell walls, entire to spinulose-dentate (especially on juvenile growth), regularly toothed, teeth terminating with mucronate to spinescent tip, apex acute or obtuse, mucronate to spinescent;

surfaces abaxially glabrate to pubescent with bluish white wax layer, often obscured by golden glandular and multiradiate hairs, adaxially yellowish green, scurfy with multiradiate hairs, later in season slightly pubescent.

Acorns

biennial;

cup turbinate to hemispheric, 7-13 mm high × 16.5-31.5 mm wide, covering 1/3-1/2 nut, outer surface light to dark reddish brown, glossy, glabrous to puberulent, inner surface light brown, glabrous, occasionally with ring of pubescence around scar, scales often tuberculate, base broad, glossy, margins strongly concave with tips tightly appressed, acute to attenuate;

nut oblong to subglobose, 12-22 × 10-21 mm, glabrous, with 1 or more rings of fine pits at apex, scar diam. 6.5-13.5 mm.

solitary or paired, rarely in 3s or 4s;

cup saucer-shaped, 4-10 mm deep × 15-40 mm wide, rims often corky and thickened, scales appressed, deeply embedded in tomentum, often appearing swollen and keeled, tuberculate;

nut ovoid, 15-30 × 10-20 mm, apex blunt, glabrous;

nut scar 4-10 mm diam.

Terminal

buds dark reddish brown, conic to ovoid, 4-7 mm, noticeably 5-angled in cross section, usually silvery- or tawny-pubescent distal to middle.

buds conic, 2-8 mm, scales brown with ciliate margins.

2n

= 24.

Quercus coccinea

Quercus chrysolepis

Phenology Flowering spring. Flowering usually in spring, occasionally in fall.
Habitat Poor soils, well-drained uplands, dry slopes, and ridges, occasionally on poorly drained sites Mountain ridges, canyons, and moist slopes
Elevation 0-1500 m (0-4900 ft) 200-2600 m (700-8500 ft)
Distribution
from FNA
AL; AR; CT; DC; DE; GA; IL; IN; KY; MA; MD; ME; MI; MO; NC; NH; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WI; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NM; NV; OR; Mexico (Baja California and Chihuahua)
[WildflowerSearch map]
[BONAP county map]
Discussion

Trees having acorns with broad, distinctly warty cups are sometimes classified as Quercus coccinea var. tuberculata Sargent.

Quercus coccinea reportedly hybridizes with Q. imbricaria, Q. ilicifolia (= Q. ×robbinsii Trelease), Q. laevis, and Q. palustris (E. J. Palmer 1948) and with Q. phellos, Q. rubra, and Q. velutina (= Q. ×fontana Laughlin).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Quercus chrysolepis is one of the most variable North American oaks. Historically, individuals with extreme variation in fruit and leaf characteristics led to the recognition of several varieties and forms; for example, shrubs with small leaves have been called Q. chrysolepis var. nana (Jepson) Jepson. Studies of quantitative and qualitative variation in these characteristics do not support the recognition of infraspecific taxa; geographic variation is apparent, however, based on populations with nearly stabilized character combinations that loosely define widespread variants. This oak is distinguished consistently from other species of the complex by the presence of multiradiate trichomes on both leaf surfaces (P. S. Manos 1993). The leaf morphology and branching habit present on juvenile growth, suckers, and shade forms may approach typical Q. palmeri. Similarly, various extreme forms often resemble other species of the complex.

Putative hybrids have been reported from narrow zones of range overlap with the three other North American species of Quercus sect. Protobalanus.

The Mendocin Indians considered the nuts of Quercus chrysolepis poisonous (D. E. Moerman 1986).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 3. FNA vol. 3.
Parent taxa Fagaceae > Quercus > sect. Lobatae Fagaceae > Quercus > sect. Protobalanus
Sibling taxa
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms Q. coccinea var. tuberculata Q. chrysolepis var. nana, Q. wilcoxii
Name authority Münchhausen: Hausvater 5(1): 254. (1770) Liebmann: Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 173. (1854)
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