Quercus coccinea |
Quercus berberidifolia |
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scarlet oak |
California scrub oak, inland scrub oak, scrub oak |
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Habit | Trees, deciduous, to 30 m; lower trunk without stubs of dead branches. | Shrubs, subevergreen, 1-2(-4) m. Bark gray, scaly. |
Bark | dark gray to dark brown, irregularly fissured with scaly ridges, inner bark orangish pink. |
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Twigs | reddish brown, (1-)2-3.5 mm diam., glabrous. |
gray, yellowish, rarely reddish, 1-3 mm diam. |
Buds | reddish brown, globose or ovoid, 2-3 mm, minutely puberulent. |
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Leaves | blade elliptic to ovate or obovate, 70-160 × 80-130 mm, base obtuse to truncate, margins with 5-9 deep lobes and 18-50 awns, lobes distally expanded, sinuses usually extending more than 1/2 distance to midrib, apex acute; surfaces abaxially glabrous except for minute axillary tufts of tomentum, adaxially glossy light green, glabrous, secondary veins raised on both surfaces. |
blade bicolored, obovate, elliptic, occasionally subrotund, planar or moderately convex, (10-)15-30 × (8-)10-20 mm, base truncate or rounded-attenuate, rarely cuneate, margins irregularly toothed and spinose, often sublobate, rarely entire, secondary veins (3-)4-7 on each side, apex broadly rounded or acute; surfaces abaxially waxy, light green or glaucous, with scattered minute, appressed, (4-)8(-10)-rayed hairs less than 0.2 mm diam. and sparse to dense yellowish, glandular hairs, adaxially glossy or dull green, glabrous or glabrate. |
Acorns | biennial; cup turbinate to hemispheric, 7-13 mm high × 16.5-31.5 mm wide, covering 1/3-1/2 nut, outer surface light to dark reddish brown, glossy, glabrous to puberulent, inner surface light brown, glabrous, occasionally with ring of pubescence around scar, scales often tuberculate, base broad, glossy, margins strongly concave with tips tightly appressed, acute to attenuate; nut oblong to subglobose, 12-22 × 10-21 mm, glabrous, with 1 or more rings of fine pits at apex, scar diam. 6.5-13.5 mm. |
solitary or paired, subsessile; cup hemispheric or turbinate, rarely shallowly cup-shaped, rim thick, 8-15 mm deep × 15-20(-22) mm wide, enclosing to 1/2 nut, scales reddish or yellowish, usually strongly, irregularly tuberculate, puberulent or canescent; nut light to dark brown, ovoid, ellipsoid, or barrel-shaped, (10-)15-30 × (8-)10-20 mm, apex rounded, glabrous at maturity. |
Cotyledons | distinct. |
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Terminal | buds dark reddish brown, conic to ovoid, 4-7 mm, noticeably 5-angled in cross section, usually silvery- or tawny-pubescent distal to middle. |
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2n | = 24. |
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Quercus coccinea |
Quercus berberidifolia |
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Phenology | Flowering spring. | Flowering spring. |
Habitat | Poor soils, well-drained uplands, dry slopes, and ridges, occasionally on poorly drained sites | Chaparral, margins of coastal sage scrub |
Elevation | 0-1500 m (0-4900 ft) | 100-1800 m (300-5900 ft) |
Distribution |
AL; AR; CT; DC; DE; GA; IL; IN; KY; MA; MD; ME; MI; MO; NC; NH; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WI; WV
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CA
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Discussion | Trees having acorns with broad, distinctly warty cups are sometimes classified as Quercus coccinea var. tuberculata Sargent. Quercus coccinea reportedly hybridizes with Q. imbricaria, Q. ilicifolia (= Q. ×robbinsii Trelease), Q. laevis, and Q. palustris (E. J. Palmer 1948) and with Q. phellos, Q. rubra, and Q. velutina (= Q. ×fontana Laughlin). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The name Quercus dumosa (see species treatment no. 72) has often been applied to this species. Quercus berberidifolia is the most common scrub oak of central and southern California, mostly at midelevations in the Coast Ranges. In central California it is replaced in drier interior habitats by Q. john-tuckeri, and south of the transverse ranges by Q. cornelius-mulleri. From Santa Barbara south, it does not descend to the low elevation coastal sites typical of Q. dumosa in the strict sense. Quercus berberidifolia hybridizes with numerous other white oaks of California. In southern California, putative hybrids with Q. john-tuckeri are noticeable in the mountains above Ventura and on the north slope of the Tehachapi Mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 3. | FNA vol. 3. |
Parent taxa | Fagaceae > Quercus > sect. Lobatae | Fagaceae > Quercus > sect. Quercus |
Sibling taxa | ||
Synonyms | Q. coccinea var. tuberculata | Q. agrifolia var. berberidifolia, Q. dumosa var. munita |
Name authority | Münchhausen: Hausvater 5(1): 254. (1770) | Liebmann: Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 172. (1854) |
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