FNA: Quercus arizonica vs. Quercus hinckleyi

	Quercus arizonica	Quercus hinckleyi
	Arizona oak, Arizona white oak	Hinckley oak
Habit	Trees, evergreen or subevergreen, small to moderate-sized trees, rarely to 18 m. Bark scaly.	Shrubs, evergreen, low, to 0.75(-1.5) m, spreading rhizomatously in thickets, intricately branched.
Bark		gray, scaly.
Twigs	yellowish, 1.5-2.5 mm diam., persistently felty-tomentose, eventually dingy gray.	light brown, pruinose, becoming waxy-glaucous in 2d season, 1-1.5 mm diam., glabrous or sparsely and minutely stellate-pubescent.
Buds	dull russet-brown, ovoid, distally subacute or rounded, 3 mm, sparsely pubescent or glabrate.	minute, subrotund, 0.5-1 mm; scales reddish brown, glabrous except for ciliate margins.
Leaves	blade elliptic or oblong to narrowly obovate or oblanceolate, planar or moderately convex, to (30-)40-80(-90) × 15-30 mm, thick and leathery, usually stiff, base cordate or rounded and weakly cordate, margins entire or coarsely toothed especially near apex, cartilaginously revolute, teeth mucronate-tipped, obscure or prominent, secondary veins ca. 7-11 on each side, branching, passing into teeth when present, apex acute to usually obtuse or broadly rounded; surfaces abaxially dull, sparsely pubescent or subtomentose with curly branched hairs, reticulate from prominent, raised secondary veins, usually glaucous where exposed, adaxially dark or bluish green, moderately lustrous, sparsely and minutely stellate-pubescent, secondary veins slightly raised or prominent within depressions or impressed.	blade subrotund or rotund, to 15 × 15 mm, thick, leathery, base cordate or auriculate, margins strongly crisped with 2-3 coarse, spinescent teeth on each side, cartilaginous-thickened secondary veins obscure, apex acute or obtuse, spine-tipped; surfaces abaxially blue-green, glaucous, glabrous, microscopically markedly papillose, adaxially blue-green, glaucous, glabrous, secondary veins slightly raised on both surfaces.
Acorns	solitary or paired, subsessile, occasionally on peduncle to 15 mm; cup hemispheric or cup-shaped, 5-10(-15) mm deep × 10-15 mm wide, enclosing ca. 1/2 nut, base rounded, margin rather coarse, scales cream to brown, broadly ovate, evenly and strongly tuberculate, tomentose, tips closely appressed; nut light brown, ovoid or oblong, 8-12 mm, nearly glabrous.	solitary, subsessile or on axillary peduncle to 4 mm; cup shallow, saucer-shaped, 1-3 mm deep × 10-15 mm wide, enclosing base of nut only, margin irregularly undulate, scales closely appressed, minute, basally tuberculate-thickened, glabrous except for thin ciliate margins; nut ovoid, 10-20 × 8-12 mm, glabrous.
Cotyledons	connate.	distinct.

	Quercus arizonica	Quercus hinckleyi
Phenology	Flowering spring.	Flowering spring.
Habitat	Oak and pinyon woodlands, margins of chaparral, arroyos	On dry desert slopes
Elevation	1300-2500(-3000) m (4300-8200(-9800) ft)	1150-1400 m (3800-4600 ft)
Distribution	AZ; NM; TX; Mexico (Chihuahua, Coahuila, Durango, and Sonora) [WildflowerSearch map] [BONAP county map]	TX; Mexico [BONAP county map]
Discussion	Some of the specimens previously referred to Quercus endemica by C. H. Muller belong here instead. Putative hybrids between Quercus arizonica and Q. grisea (= Q. ×organensis Trelease) are problematic in local areas of contact from southeastern Arizona to western Texas. These intermediates tend to have narrower leaves than Q. arizonica, with moderately reticulate patterns of venation, and more densely hairy leaves. Quercus arizonica and Q. grisea are amply distinct elsewhere, including large areas in northern Mexico, and they appear to be more closely related to other species than to one another (e.g., Q. arizonica with Q. oblongifolia and Q. laeta Liebmann, and Q. grisea with Q. mohriana and Q. microphylla Née). Thus, Q. arizonica and Q. grisea are best treated as distinct species that hybridize, and not as conspecific populations. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Of conservation concern. This species is known only from two sites in the United States, El Solitario and near Shafter, Texas. The Shafter population includes some individuals with characteristics that suggest hybridization with Quercus pungens Liebmann. These plants are larger, with more pubescent twigs and leaves, and hemispheric acorn cups to 10 mm deep. Such plants have recently been collected in adjacent Mexico. Fossil evidence from packrat middens indicates Q. hinckleyi probably had a broader distribution and was a dominant shrub between 19,000 and 9500 years ago. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Fagaceae > Quercus > sect. Quercus	Fagaceae > Quercus > sect. Quercus
Sibling taxa	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii	Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii
Synonyms	Q. sacame
Name authority	Sargent: Gard. & Forest 8: 92. (1895)	C. H. Muller: Contr. Texas Res. Found., Bot. Stud. 1: 40. (1951)
Web links	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Quercus hinckleyi

Arizona oak, Arizona white oak

Hinckley oak

Habit

Trees, evergreen or subevergreen, small to moderate-sized trees, rarely to 18 m. Bark scaly.

Shrubs, evergreen, low, to 0.75(-1.5) m, spreading rhizomatously in thickets, intricately branched.

Bark

gray, scaly.

Twigs

yellowish, 1.5-2.5 mm diam., persistently felty-tomentose, eventually dingy gray.

light brown, pruinose, becoming waxy-glaucous in 2d season, 1-1.5 mm diam., glabrous or sparsely and minutely stellate-pubescent.

Buds

dull russet-brown, ovoid, distally subacute or rounded, 3 mm, sparsely pubescent or glabrate.

minute, subrotund, 0.5-1 mm;

scales reddish brown, glabrous except for ciliate margins.

Leaves

blade elliptic or oblong to narrowly obovate or oblanceolate, planar or moderately convex, to (30-)40-80(-90) × 15-30 mm, thick and leathery, usually stiff, base cordate or rounded and weakly cordate, margins entire or coarsely toothed especially near apex, cartilaginously revolute, teeth mucronate-tipped, obscure or prominent, secondary veins ca. 7-11 on each side, branching, passing into teeth when present, apex acute to usually obtuse or broadly rounded;

surfaces abaxially dull, sparsely pubescent or subtomentose with curly branched hairs, reticulate from prominent, raised secondary veins, usually glaucous where exposed, adaxially dark or bluish green, moderately lustrous, sparsely and minutely stellate-pubescent, secondary veins slightly raised or prominent within depressions or impressed.

blade subrotund or rotund, to 15 × 15 mm, thick, leathery, base cordate or auriculate, margins strongly crisped with 2-3 coarse, spinescent teeth on each side, cartilaginous-thickened secondary veins obscure, apex acute or obtuse, spine-tipped;

surfaces abaxially blue-green, glaucous, glabrous, microscopically markedly papillose, adaxially blue-green, glaucous, glabrous, secondary veins slightly raised on both surfaces.

Acorns

solitary or paired, subsessile, occasionally on peduncle to 15 mm;

cup hemispheric or cup-shaped, 5-10(-15) mm deep × 10-15 mm wide, enclosing ca. 1/2 nut, base rounded, margin rather coarse, scales cream to brown, broadly ovate, evenly and strongly tuberculate, tomentose, tips closely appressed;

nut light brown, ovoid or oblong, 8-12 mm, nearly glabrous.

solitary, subsessile or on axillary peduncle to 4 mm;

cup shallow, saucer-shaped, 1-3 mm deep × 10-15 mm wide, enclosing base of nut only, margin irregularly undulate, scales closely appressed, minute, basally tuberculate-thickened, glabrous except for thin ciliate margins;

nut ovoid, 10-20 × 8-12 mm, glabrous.

Cotyledons

connate.

distinct.

Quercus arizonica

Quercus hinckleyi

Phenology

Flowering spring.

Habitat

Oak and pinyon woodlands, margins of chaparral, arroyos

On dry desert slopes

Elevation

1300-2500(-3000) m (4300-8200(-9800) ft)

1150-1400 m (3800-4600 ft)

Distribution

AZ; NM; TX; Mexico (Chihuahua, Coahuila, Durango, and Sonora)

[WildflowerSearch map]

[BONAP county map]

TX; Mexico

[BONAP county map]

Discussion

Some of the specimens previously referred to Quercus endemica by C. H. Muller belong here instead.

Putative hybrids between Quercus arizonica and Q. grisea (= Q. ×organensis Trelease) are problematic in local areas of contact from southeastern Arizona to western Texas. These intermediates tend to have narrower leaves than Q. arizonica, with moderately reticulate patterns of venation, and more densely hairy leaves. Quercus arizonica and Q. grisea are amply distinct elsewhere, including large areas in northern Mexico, and they appear to be more closely related to other species than to one another (e.g., Q. arizonica with Q. oblongifolia and Q. laeta Liebmann, and Q. grisea with Q. mohriana and Q. microphylla Née). Thus, Q. arizonica and Q. grisea are best treated as distinct species that hybridize, and not as conspecific populations.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

This species is known only from two sites in the United States, El Solitario and near Shafter, Texas. The Shafter population includes some individuals with characteristics that suggest hybridization with Quercus pungens Liebmann. These plants are larger, with more pubescent twigs and leaves, and hemispheric acorn cups to 10 mm deep. Such plants have recently been collected in adjacent Mexico. Fossil evidence from packrat middens indicates Q. hinckleyi probably had a broader distribution and was a dominant shrub between 19,000 and 9500 years ago.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Fagaceae > Quercus > sect. Quercus

Sibling taxa

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hinckleyi, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Q. acerifolia, Q. agrifolia, Q. ajoensis, Q. alba, Q. arizonica, Q. arkansana, Q. austrina, Q. berberidifolia, Q. bicolor, Q. boyntonii, Q. buckleyi, Q. carmenensis, Q. chapmanii, Q. chihuahuensis, Q. chrysolepis, Q. coccinea, Q. cornelius-mulleri, Q. depressipes, Q. douglasii, Q. dumosa, Q. durata, Q. ellipsoidalis, Q. emoryi, Q. engelmannii, Q. falcata, Q. fusiformis, Q. gambelii, Q. garryana, Q. geminata, Q. georgiana, Q. graciliformis, Q. gravesii, Q. grisea, Q. havardii, Q. hemisphaerica, Q. hypoleucoides, Q. ilicifolia, Q. imbricaria, Q. incana, Q. inopina, Q. intricata, Q. john-tuckeri, Q. kelloggii, Q. laceyi, Q. laevis, Q. laurifolia, Q. lobata, Q. lyrata, Q. macrocarpa, Q. margarettae, Q. marilandica, Q. michauxii, Q. minima, Q. mohriana, Q. montana, Q. muehlenbergii, Q. myrtifolia, Q. nigra, Q. oblongifolia, Q. oglethorpensis, Q. pacifica, Q. pagoda, Q. palmeri, Q. palustris, Q. phellos, Q. polymorpha, Q. prinoides, Q. pumila, Q. pungens, Q. robur, Q. robusta, Q. rubra, Q. rugosa, Q. sadleriana, Q. shumardii, Q. similis, Q. sinuata, Q. stellata, Q. tardifolia, Q. texana, Q. tomentella, Q. toumeyi, Q. turbinella, Q. vacciniifolia, Q. vaseyana, Q. velutina, Q. viminea, Q. virginiana, Q. wislizenii

Synonyms

Q. sacame

Name authority

Sargent: Gard. & Forest 8: 92. (1895)

C. H. Muller: Contr. Texas Res. Found., Bot. Stud. 1: 40. (1951)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Flora of North America species comparison

Quercus arizonica

Quercus hinckleyi

Quercus arizonica

Quercus hinckleyi