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tuberous desert-chicory

desert-chicory

Habit Perennials (possibly flowering first year), 5–30(–45+) cm (roots or rootstocks producing tuberiform swellings 1–15 cm below soil surface). Annuals or perennials, 5–100+ cm; taprooted or rhizomatous (roots producing tuberiform swellings in P. grandiflorus).
Stems

usually scapiform, usually branching from bases, proximally glabrous or sparsely to densely pilosulous.

usually 1, sometimes 2–5+, erect, unbranched or branched proximally and/or distally, glabrous or pilosulous.

Leaves

basal or basal and cauline;

basal ± petiolate, distal usually sessile;

blades oblong, elliptic, or ovate to lanceolate or linear, margins entire or dentate to pinnately lobed (faces usually glabrous, sometimes pilosulous near margins).

Cauline leaves

0(–3), proximal mostly lanceolate, margins usually pinnately lobed, distal linear-filiform, margins entire.

Peduncles

not inflated distally, sometimes bracteate.

Involucres

cylindric to turbinate, 17–25 mm.

cylindric, 4–5[–8+] mm diam.

Receptacles

± convex, pitted, glabrous, epaleate.

Florets

40–60+;

anthers 4.5–5 mm (pollen equatorial diameters 46–52 µm).

(20–)30–150+;

corollas yellow to whitish.

Phyllaries

13–22.

8–21+ in ± 2 series (reflexed in fruit), linear, equal, margins often scarious, apices acute (often thickened or bearing keel-like flaps near tips).

Calyculi

bractlets 8 in 1(–2) series, linear to subulate, 3–8 mm.

of 3–13+, deltate to subulate or filiform bractlets.

Heads

borne singly or 2–3 in loose, corymbiform arrays.

borne singly or in loose, corymbiform arrays.

Cypselae

bodies stramineous, 4–5 mm, beaks 6–7 mm;

pappi 10–12 mm.

reddish brown to stramineous, bodies ± fusiform, beaks (± concolorous with bodies) ± filiform, fragile, grooves (or broad ribs) 5, faces transversely rugulose, glabrous;

pappi (borne on discs at tips of beaks) persistent, double: outer coroniform (of whitish, relatively short, spreading, sometimes curly, hairs), inner of 80–120+, rufous to stramineous, subequal, barbellulate bristles in 2–3+ series.

x

= 6.

2n

= 24.

Pyrrhopappus grandiflorus

Pyrrhopappus

Phenology Flowering Apr–Jun.
Habitat Disturbed sites, calcareous, loamy, or sandy soils
Elevation 100–800 m (300–2600 ft)
Distribution
from FNA
KS; OK; TX
[WildflowerSearch map]
[BONAP county map]
from USDA
North America; Mexico
[BONAP county map]
Discussion

Pyrrhopappus grandiflorus has been reported from Arkansas and New Mexico; I have seen no specimens of it from those states.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 1, 4, or 5 (4 in the flora).

Some plants from Mexico that have been called Pyrrhopappus multicaulis de Candolle (type from Mexico) may be distinct at species rank from P. pauciflorus (see R. McVaugh 1984).

Almost all botanists who have dealt with the biology or floristics of pyrrhopappuses have remarked similarities among the taxa and/or difficulties with identification of some specimens. Some botanists have noted that “interspecific” hybridizations are common (D. K. Northington 1974; B. L. Turner and K. J. Kim 1990; and works cited therein).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Perennials (roots or rootstocks producing tuberiform swellings 1–15 cm below soil surface); stems usually scapiform; cauline leaves usually 0, sometimes 1–3; anthers 4.5–5 mm.
P. grandiflorus
1. Annuals or perennials (not producing tuberiform swellings); stems rarely scapiform; cauline leaves usually 3–9+, seldom 0–2; anthers 2.5–4 mm
→ 2
2. Stems usually sparsely to densely pilosulous proximally, sometimes glabrous; blades of distal cauline leaves usually pinnately (3–)5–7(–9+)-lobed
P. pauciflorus
2. Stems usually glabrous proximally, sometimes pilosulous; blades of distal cauline leaves usually entire or with 1–2 lobes near bases, sometimes pinnately 3–5(–7+)-lobed
→ 3
3. Perennials; involucres 12–15(–17+) mm; florets 30–60+; cypselae stramineous; pappi 6–7 mm
P. rothrockii
3. Annuals; involucres 17–24 mm; florets 50–150+; cypselae reddish brown; pappi 9–10 mm
P. carolinianus
Source FNA vol. 19, p. 377. FNA vol. 19, p. 376. Author: John L. Strother.
Parent taxa Asteraceae > tribe Cichorieae > Pyrrhopappus Asteraceae > tribe Cichorieae
Sibling taxa
P. carolinianus, P. pauciflorus, P. rothrockii
Subordinate taxa
P. carolinianus, P. grandiflorus, P. pauciflorus, P. rothrockii
Synonyms Barkhausia grandiflora
Name authority (Nuttall) Nuttall: Trans. Amer. Philos. Soc., n. s. 7: 430. (1841) de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 7: 144. (1838)
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