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Nuttall alkaligrass, Nuttall's alkali grass

Howell's alkali grass

Habit Plants perennial; cespitose, not mat-forming. Plants perennial; cespitose, not mat-forming.
Culms

10-100 cm, usually erect.

7-40 cm, erect to ascending.

Leaves

either concentrated at the base or distributed along the culms;

ligules 1-3 mm, obtuse, usually entire, sometimes slightly erose;

blades 1-4 mm wide, flat to involute.

Panicles

5-30 cm, compact to diffuse at maturity, usually distinctly exserted from the sheaths, lower branches usually erect to diverging, occasionally descending, spikelet-bearing from the base or on the distal 2/3;

pedicels scabrous, lacking tumid epidermal cells.

2-13 cm, compact to diffuse at maturity, lower branches erect to descending, spikelet-bearing from near the base or confined to the distal 1/2;

pedicels smooth or with a few scattered scabrules, sometimes with tumid epidermal cells.

Spikelets

3.5-9 mm, with 2-7 florets.

3-8 mm, with (1)2-5 florets.

Glumes

rounded over the back, veins obscure, apices acute to obtuse;

lower glumes 0.5-1.5 mm, usually less than 1/2 as long as the adjacent lemmas;

upper glumes 1-2.8 mm;

rachilla internodes slightly and gradually broadened to the point of attachment with the lemmas;

calluses with a few hairs;

lemmas (2)2.2-3(3.5) mm, herbaceous, glabrous or sparsely hairy on the proximal 1/2, principally along the veins, backs rounded, 5-veined, veins obscure, not extending to the margins, smooth distally, lateral margins inrolled or not, apical margins uniformly and densely scabrous, apices acute to obtuse, entire;

palea veins glabrous, short-ciliate, or with a few long hairs proximally, smooth or scabrous distally;

anthers 0.6-2 mm.

rounded or weakly keeled over the back, veins obscure, apices acute to obtuse;

lower glumes 0.8-1.9 mm;

upper glumes 1.7-2.5 mm;

calluses with a few hairs;

lemmas 2.4-3.3 mm, herbaceous, glabrous or with a few hairs near the base, principally along the veins, backs rounded or weakly keeled distally, 5-veined, veins obscure, not extending to the margins, apical margins uniformly and densely scabrous, apices acute to obtuse, entire;

palea veins smooth and glabrous proximally, smooth or scabrous distally;

anthers 1.5-2 mm.

Ligules

1.5-2.7 mm, obtuse, entire or minutely and irregularly serrate;

blades involute, 1.4-2.2 mm wide when flattened.

2n

= 28, 42, 56.

= unknown.

Puccinellia nuttalliana

Puccinellia howellii

Distribution
from FNA
AK; AZ; CA; CO; ID; KS; ME; MN; MT; ND; NE; NM; NV; NY; OR; SD; UT; VT; WA; WI; WY; AB; BC; MB; NT; NU; ON; PE; QC; SK; YT; Greenland
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Puccinellia nuttalliana is a widespread and variable species, restricted to the Flora region. It grows principally in the interior, but is also found in coastal settings, where it is difficult to distinguish from P. nutkaensis. Northern, primarily boreal or southern arctic populations with relatively short lemmas and anthers (2-2.8 mm and 0.6-0.9 mm, respectively), and with a few long hairs on the lower palea veins, have sometimes been recognized as P. borealis Swallen.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Puccinellia howellii is known only from the type locality in Shasta County, California, where it is a dominant element of the vegetation associated with a group of three mineralized seeps. Isozyme profiles suggest that it is a polyploid.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 475. FNA vol. 24, p. 475.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Puccinellia Poaceae > subfam. Pooideae > tribe Poeae > Puccinellia
Sibling taxa
P. andersonii, P. angustata, P. arctica, P. bruggemannii, P. distans, P. fasciculata, P. groenlandica, P. howellii, P. lemmonii, P. maritima, P. nutkaensis, P. parishii, P. phryganodes, P. pumila, P. rupestris, P. simplex, P. tenella, P. vaginata, P. vahliana, P. wrightii
P. andersonii, P. angustata, P. arctica, P. bruggemannii, P. distans, P. fasciculata, P. groenlandica, P. lemmonii, P. maritima, P. nutkaensis, P. nuttalliana, P. parishii, P. phryganodes, P. pumila, P. rupestris, P. simplex, P. tenella, P. vaginata, P. vahliana, P. wrightii
Synonyms P. rosenkrantzii, P. porsildii, P. interior, P. deschampsioides, P. cusickii, P. borealis, P. airoides
Name authority (Schult.) Hitchc. J.I. Davis
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