Ptychostomum creberrimum |
Ptychostomum warneum |
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Habit | Plants in dense or open turfs, green or yellow-green. | Plants in dense or open turfs, red, green, or yellow-green. |
Stems | 1–2(–3) cm, comose, innovations comose or somewhat elongate and evenly foliate; often strongly radiculose. |
0.5–2(–3) cm, fertile stems comose, innovations evenly foliate. |
Leaves | green, twisted to contorted when dry, ovate-lanceolate, flat to weakly concave, (1–)2–3(–3.5) mm, somewhat enlarged toward stem apex; base weakly decurrent; margins revolute to mid leaf or beyond, limbidium strong, in 2 or 3 rows, yellowish; apex acuminate; costa long-excurrent, awn ± smooth; proximal laminal cells 3–4:1, same width or sometimes wider than more distal cells; medial and distal cells rhomboidal, 14–22 µm wide, 3–4:1, walls usually thin to firm, never oblique to costa. |
yellow-green to red, crowded, strongly contorted to shrunken when dry, ovate-lanceolate, flat, 0.5–2 mm, often gradually enlarged toward stem apex; base usually green, not decurrent; margins revolute proximally, limbidium strong, in 2 or 3 rows; apex acuminate; costa short-excurrent, awn slender; proximal laminal cells rectangular, 3–4:1; medial and distal cells 18–22 µm wide, 2–3:1, walls thin. |
Seta | 1–2(–3) cm. |
purple or red, 2–4(–5) cm, stout, straight to somewhat flexuose. |
Sexual condition | synoicous. |
usually polyoicous, sometimes autoicous. |
Capsule | brown, elongate-pyriform, symmetric, 2–4 mm, mouth yellow; operculum conic, apiculate; peristome well developed; exostome teeth yellow basally, hyaline distally, lamellae usually straight mid tooth, pores absent along mid line; endostome not adherent to exostome, basal membrane high, 1/2 exostome height, segments with broadly ovate perforations, cilia long, appendiculate. |
yellow-brown or brown, pyriform, symmetric, 2–4 mm, mouth yellow; hypophysis slender; operculum long-conic; peristome reduced; exostome teeth yellow or brown basally, pale yellow to hyaline distally, lamellae straight, pores absent near base along mid line; endostome weakly adherent to exostome, basal membrane 1/2 exostome height, segments narrowly perforate, cilia absent or rudimentary. |
Spores | 10–14(–16) µm, finely papillose, pale brown-yellow or green. |
with size variable in same capsule, (32–)36–48(–50) µm, pale green or yellow. |
Specialized | asexual reproduction absent. |
asexual reproduction absent. |
Ptychostomum creberrimum |
Ptychostomum warneum |
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Phenology | Capsules mature May–Sep. | Capsules mature Jul–Aug. |
Habitat | Damp to dry soil, soil over rock | Damp soil |
Elevation | low to high elevations (0-3500 m) (low to high elevations (0-11500 ft)) | low elevations (0-50 m) (low elevations (0-200 ft)) |
Distribution |
AK; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; VT; WA; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; YT; Pacific Islands (New Zealand); Greenland; Eurasia; s South America; Australia |
QC; Greenland; n Eurasia (Himalayan Nepal) |
Discussion | Ptychostomum creberrimum is one of the most widespread and common species in the genus, occurring throughout the Northern Hemisphere and disjunct in the Southern Hemisphere. The species can be distinguished from the closely related P. lonchocaulon by its strictly synoicous sexual condition and narrower yellowish leaf border. It is similar also to P. pallescens, which is autoicous and has larger spores and elongate-ovate segment perforations. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Ptychostomum warneum is an arctic-boreal species that is distinguished from the related P. calophyllum by its pyriform brown capsule with a slender neck, generally larger spores, and acuminate leaves. The stems are sparsely radiculose; the costal awns are smooth. The species is known from only one location in Quebec, in the Mingan Islands. Capsules are needed for identification. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 28, p. 159. | FNA vol. 28, p. 174. |
Parent taxa | Bryaceae > Ptychostomum > subg. Cladodium | Bryaceae > Ptychostomum > subg. Ptychostomum |
Sibling taxa | ||
Synonyms | Bryum creberrimum, B. lisae var. cuspidatum | Mnium caespiticium var. warneum, Bryum warneum |
Name authority | (Taylor) J. R. Spence & H. P. Ramsay: Phytologia 87: 23. (2005) | (Röhling) J. R. Spence: Phytologia 87: 22. (2005) |
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