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Habit Plants in dense or open turfs, green or yellow-green. Plants in dense turfs, dark green.
Stems

1–2(–3) cm, comose, innovations comose or somewhat elongate and evenly foliate; often strongly radiculose.

1–2 cm, comose, innovations comose or somewhat elongate and evenly foliate;

not strongly radiculose.

Leaves

green, twisted to contorted when dry, ovate-lanceolate, flat to weakly concave, (1–)2–3(–3.5) mm, somewhat enlarged toward stem apex;

base weakly decurrent;

margins revolute to mid leaf or beyond, limbidium strong, in 2 or 3 rows, yellowish;

apex acuminate;

costa long-excurrent, awn ± smooth;

proximal laminal cells 3–4:1, same width or sometimes wider than more distal cells;

medial and distal cells rhomboidal, 14–22 µm wide, 3–4:1, walls usually thin to firm, never oblique to costa.

green, twisted to contorted when dry, oblong or ovate, concave, (0.5–)1–2 mm, not much enlarged toward stem apex;

base not decurrent;

margins plane or revolute to mid leaf, limbidium somewhat indistinct, in 1 or 2 rows;

apex acute, sometimes apiculate;

costa percurrent in proximal leaves to sometimes short-excurrent in distal leaves, awn denticulate;

proximal laminal cells 3–4:1, same width or sometimes wider than more distal cells;

medial and distal cells rhomboidal, 12–16 µm wide, 3–4:1, walls thin.

Seta

1–2(–3) cm.

2–3 cm.

Sexual condition

synoicous.

autoicous.

Capsule

brown, elongate-pyriform, symmetric, 2–4 mm, mouth yellow;

operculum conic, apiculate;

peristome well developed;

exostome teeth yellow basally, hyaline distally, lamellae usually straight mid tooth, pores absent along mid line;

endostome not adherent to exostome, basal membrane high, 1/2 exostome height, segments with broadly ovate perforations, cilia long, appendiculate.

brown, pyriform, symmetric, 2–3 mm, mouth yellow;

operculum short-conic, apiculate;

peristome reduced;

exostome teeth yellow basally, hyaline distally, lamellae usually straight mid tooth, pores absent along mid line;

endostome variable, not adherent to exostome, basal membrane low, segments with narrowly ovate perforations, sometimes reduced, cilia variable, absent or occasionally present and well developed, appendiculate or nodose.

Spores

10–14(–16) µm, finely papillose, pale brown-yellow or green.

(28–)30–40 µm, finely papillose, green.

Specialized

asexual reproduction absent.

asexual reproduction by leaf axil gemmae, green or green-brown.

Ptychostomum creberrimum

Ptychostomum reedii

Phenology Capsules mature May–Sep. Capsules mature Jun–Aug.
Habitat Damp to dry soil, soil over rock Soil over serpentine rock, in serpentine rock crevices
Elevation low to high elevations (0-3500 m) (low to high elevations (0-11500 ft)) low elevations (0-100 m) (low elevations (0-300 ft))
Distribution
from FNA
AK; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; VT; WA; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; YT; Pacific Islands (New Zealand); Greenland; Eurasia; s South America; Australia
from FNA
DE; MD
Discussion

Ptychostomum creberrimum is one of the most widespread and common species in the genus, occurring throughout the Northern Hemisphere and disjunct in the Southern Hemisphere. The species can be distinguished from the closely related P. lonchocaulon by its strictly synoicous sexual condition and narrower yellowish leaf border. It is similar also to P. pallescens, which is autoicous and has larger spores and elongate-ovate segment perforations.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Ptychostomum reedii is a highly distinctive and very rare local endemic, not likely to be confused with any other species in eastern North America. The costal awns are slender. The setae are brown, unlike the red or red-brown setae of other species in sect. Cladodium. Plants from Delaware have abundant filiform gemmae in the leaf axils.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 159. FNA vol. 28, p. 164.
Parent taxa Bryaceae > Ptychostomum > subg. Cladodium Bryaceae > Ptychostomum > subg. Cladodium
Sibling taxa
P. acutiforme, P. archangelicum, P. arcticum, P. bimum, P. calophyllum, P. cernuum, P. cryophilum, P. cyclophyllum, P. inclinatum, P. intermedium, P. knowltonii, P. lonchocaulon, P. longisetum, P. marratii, P. meesioides, P. neodamense, P. nitidulum, P. pallens, P. pallescens, P. pendulum, P. pseudotriquetrum, P. reedii, P. rutilans, P. salinum, P. schleicheri, P. subneodamense, P. turbinatum, P. warneum, P. weigelii, P. wrightii
P. acutiforme, P. archangelicum, P. arcticum, P. bimum, P. calophyllum, P. cernuum, P. creberrimum, P. cryophilum, P. cyclophyllum, P. inclinatum, P. intermedium, P. knowltonii, P. lonchocaulon, P. longisetum, P. marratii, P. meesioides, P. neodamense, P. nitidulum, P. pallens, P. pallescens, P. pendulum, P. pseudotriquetrum, P. rutilans, P. salinum, P. schleicheri, P. subneodamense, P. turbinatum, P. warneum, P. weigelii, P. wrightii
Synonyms Bryum creberrimum, B. lisae var. cuspidatum Bryum reedii
Name authority (Taylor) J. R. Spence & H. P. Ramsay: Phytologia 87: 23. (2005) (H. Robinson) J. R. Spence: Phytologia 87: 22. (2005)
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