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incurved grizzled moss, ptychomitrium moss

ptychomitrium moss

Habit Plants small, cespitose, glossy, dark green. Plants small to robust, tufted or caspitose, dark green to blackish.
Stems

erect, to 0.5 cm.

erect or repent.

Leaves

crispate when dry, oblong-lanceolate, 2 mm;

margins entire distally, erect proximally;

apex cucullate to subcucullate.

erect to crispate when dry, margins entire to serrulate or serrate;

medial cells smooth or slightly papillose.

Seta

1 per perichaetium, 2–3 mm.

straight.

Capsule

ovoid, 0.75–1 mm, smooth or wrinkled when dry;

peristome teeth perforate but mostly not divided, densely papillose.

ovoid to cylindric, symmetric or slightly curved, smooth to wrinkled or ribbed when dry.

Calyptra

lobes about one third length of calyptra.

mitrate, more or less plicate, lobed proximally.

Spores

smooth to papillose.

Specialized

asexual reproduction rare, short-uniseriate or branched gemmae, several cells long, on branched axillary filaments.

asexual reproduction rare, by 1-seriate gemmae on branched axillary filaments.

Ptychomitrium incurvum

Ptychomitrium

Phenology Capsules mature Sep–Jun.
Habitat Acidic and calcareous rocks, soil, very rarely tree bark, open forests
Elevation low to moderate elevations (0-1400 m) (low to moderate elevations (0-4600 ft))
Distribution
from FNA
AL; AR; CT; DC; DE; FL; GA; IL; IN; KS; KY; LA; MD; MI; MO; MS; NC; NJ; NY; OH; OK; PA; SC; TN; TX; VA; WV; South America (Ecuador); Europe
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Nearly worldwide; mostly in temperate regions
Discussion

The small dark green plants of Ptychomitrium incurvum are unmistakable growing on rock, with their glossy leaves tightly crispate when dry. The leaves are shorter than in P. sinense and are straight when wet, not somewhat falcate at the tips as is common in P. sinense. Sporophytes are very common in this moss and the old sporophytes persist for a long time. Sterile colonies of P. incurvum can be very reminiscent of Weissia controversa, but Ptychomitrium is more glossy; its leaves have smooth cells and without the strongly involute margins of Weissia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 40–50 (5 in the flora).

Excluded Species:

Glyphomitrium canadense Mitten

This taxon was included for western Canada by G. N. Jones (1933), but was excluded from North America by H. A. Crum (1972) and L. E. Anderson et al. (1990). Crum considered Glyphomitrium canadense to be a synonym of the British G. daviesii (Withering) Bridel and suggested that the specimen on which Mitten based the name came from Great Britain.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves coarsely serrate distally
→ 2
1. Leaves entire or obscurely irregularly serrulate distally
→ 3
2. Leaves slenderly long-acuminate, 4-6 mm; basal leaf margins broadly recurved on one or both sides; calyptra deeply lobed proximally, lobes half length of calyptra.
P. gardneri
2. Leaves broadly acuminate, 3-4 mm; basal leaf margins plane and erect or irregularly narrowly recurved on one side proximally; calyptra shallowly lobed proximally, lobes less than 1/2 length of calyptra.
P. serratum
3. Plants dull, mostly corticolous; leaves straight or slightly contorted but not crispate when dry, margins mostly obscurely serrulate distally.
P. drummondii
3. Plants glossy, mostly on rock; leaves crispate when dry, margins entire
→ 4
4. Longest leaves mostly 2.5-4 mm.
P. sinense
4. Longest leaves mostly 2 mm.
P. incurvum
Source FNA vol. 27, p. 309. FNA vol. 27, p. 307.
Parent taxa Ptychomitriaceae > Ptychomitrium Ptychomitriaceae
Sibling taxa
P. drummondii, P. gardneri, P. serratum, P. sinense
Subordinate taxa
P. drummondii, P. gardneri, P. incurvum, P. serratum, P. sinense
Synonyms Weissia incurva
Name authority (Schwägrichen) Spruce: Ann. Mag. Nat. Hist., ser. 2, 3: 487. (1849) Fürnrohr: Flora 12(Ergänzungsbl.): 19. (1829)
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