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Chinese brake, Huguenot fern, saw-leaf bracken, spider brake, spider fern

brake, brake fern

Habit Plants terrestrial or on rock.
Stems

slender, short-creeping, densely scaly;

scales dark reddish brown to chestnut brown.

erect or creeping, branched;

scales pale brown to black, concolored, elongate, margins entire.

Leaves

clustered, 1–6 dm.

monomorphic, clustered or closely spaced, 1–20 dm.

Petiole

pale or brownish, 5–30 cm, scaly proximally, otherwise glabrous.

straw-colored, green, brownish red to purple black, longitudinally ridged, 2–3-grooved adaxially, scaly at base, glabrous or scaly distally, with 1 (less often 2 or more) vascular bundle.

Blade

oblong to oblanceolate, irregularly and pedately divided proximally (as in Pteris cretica) and pinnately divided distally, 10–35 × 13–25 cm;

rachis slightly and evenly winged, wing constricted above each pinna pair.

oblong to lanceolate to deltate, 1–4-pinnate, herbaceous to leathery, abaxially and adaxially glabrous or sometimes pubescent or scaly, adaxially dull, not striate;

rachis straight.

Ultimate segments

of blade sessile to short-stalked, linear to oblong-lanceolate, 1.5–8 mm wide;

base truncate or narrowed to stalk, stalk when present green, not lustrous;

margins plane or reflexed to form false indusia.

Pinnae

3–7 pairs, widely spaced, distal pinnae simple, adnate and decurrent to rachis;

pinnae remaining green through winter, not articulate to rachis, lanceolate to linear;

sterile pinnae wider than fertile pinnae (to ca. 1.2 cm), margins irregularly serrate to serrulate;

fertile pinnae mostly less than 5 mm, margins entire to serrate at apex;

adaxial costae with sparse, septate hairs;

proximal pinnae with 1–4 elongate basal segments.

Veins

free, simple or forked.

in leaves conspicuous, free (except in sori) and forking well above base of segment, or highly anastomosing.

False indusia

pale, scarious, covering sori.

Sori

narrow, blade tissue exposed abaxially.

Sporangia

intramarginal, sori usually continuous except at pinna or segment apex and sinuses, paraphyses present.

Spores

brown, trilete, tetrahedral, rugate and/or tuberculate, usually with prominent equatorial flange.

x

= 29.

2n

= 116.

Pteris multifida

Pteris

Habitat Terrestrial or on rock in disturbed areas in circumneutral soils; primarily coastal plain.
Distribution
from FNA
AL; AR; FL; GA; IL; IN; KY; LA; MD; MS; NC; NY; SC; TX; West Indies; South America in Argentina; Brazil; native to Asia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from USDA
Worldwide; warm and tropical regions
[BONAP county map]
Discussion

Pteris multifida is found on old shady walls and masonry around cemeteries, dumps, and towns. It may no longer occur in Indiana. Juveniles of Pteris multifida may key to Pteris cretica.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species ca. 300 (5 species and 1 hybrid in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Veins in leaves anastomosing except sometimes near margins of ultimate segments.
P. tripartita
1. Veins in leaves entirely free.
→ 2
2. Leaves not strictly 1-pinnate, at least proximal pinnae pinnatifid-lobed or variously forked or divided.
→ 3
2. Leaves strictly 1-pinnate, pinnae not lobed or divided.
→ 4
3. Pinnae of mature leaves decurrent to relatively broad-winged rachis in at least distal 1/2 of leaf.
P. multifida
3. Pinnae of mature leaves not decurrent to relatively broad-winged rachis or only terminal pinna decurrent on rachis.
P. cretica
4. Petioles and often also rachises densely scaly, scales light to reddish, often grading into hairs on abaxial costae; pinnae appearing not articulate to rachis, apices long-attenuate or sharply acute; sori narrow, with most of abaxial blade surface exposed.
P. vittata
4. Petioles often sparsely scaly or scaly only proximally, scales dark brown to nearly black, scales absent or few on rachises, abaxial costae with or without hairs; pinnae appearing articulate to rachis, apices acute; sori broad, little or no abaxial blade tissue exposed.
P. bahamensis
Source FNA vol. 2, p. 134. FNA vol. 2. Author: Clifton E. Nauman.
Parent taxa Pteridaceae > Pteris Pteridaceae
Sibling taxa
P. bahamensis, P. cretica, P. tripartita, P. vittata
Subordinate taxa
P. bahamensis, P. cretica, P. multifida, P. tripartita, P. vittata
Synonyms Pycnodoria multifida
Name authority Poiret: in Lamarck et al., Encycl. 5: 714. (1804) Linnaeus: Sp. Pl. 2: 1073. 1753; Gen. Pl. ed 5, 484. (1754)
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