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dalea, indigo-bush

dotted dalea, Nevada dalea

Habit Shrubs, subshrubs, or trees, armed or unarmed, sterile shoots sometimes sharp-tipped. Shrubs.
Stems

erect, usually pubescent, rarely glabrate or glabrous, glandular nearly throughout, gland-dotted when young.

4–10(–15) dm, densely gland-dotted when young, silky-pilosulous, hairs retrorse.

Branches

divaricate;

sterile shoots almost sharp-tipped.

Leaves

alternate, odd-pinnate or unifoliolate;

stipules present, caducous, usually triangular to subulate or linear, rarely obovate;

petiolate;

leaflets 1–17(or 19), stipels absent, blade margins entire or gland-crenulate, surfaces usually glandular-punctate abaxially, pubescent.

pinnate, 0.7–2.7 cm;

leaflets 7–13, blades obovate to suborbiculate, 1–4.5 mm, terminal leaflet shorter than laterals, surfaces with submarginal and apical glands abaxially, glands visible, pubescent.

Racemes

dense to relatively loose;

rachis not spine-tipped in anthesis, 0.3–3 cm;

bracts lanceolate or linear-lanceolate, 1.6–6.5 mm;

bracteoles absent.

Inflorescences

1–20+-flowered, terminal and axillary (sometimes leaf-opposed), usually racemes, rarely spikes;

bracts present, caducous;

bracteoles 0 or 2, at apex of pedicel or base of calyx.

Flowers

papilionaceous;

calyx campanulate, lobes 5, unequal;

ribs 10, not anastomosing into closed arches distally;

corolla usually blue, blue-purple, pink-purple, magenta-purple, violet-purple, violet, or bicolored, sometimes with yellow eye, rarely white;

banner blade differentiated from claw, reflexed less than 90°, claw shorter;

wings not adnate to keel, blades oblique basally, claws shorter, linear;

keel not strongly twisted, blades narrowly overlapping and adherent, oblique basally, blunt-tipped, claws linear;

stamens 10, monadelphous, equal or alternately short and long;

anthers dorsifixed;

style glabrous or pilosulous.

calyx 3.9–5.3(–8.3) mm, pilosulous throughout or tube glabrous, tube 1.9–2.8(–3) mm, ribs prominent, intervals each with 1 row of 2–6 glands, lobes triangular-lanceolate to lanceolate, abaxial lobe longer;

corolla pink-purple, banner with yellow eye;

banner 4.6–6.1(–6.6) mm, ovate-cordate, ovate, base cordate, apex emarginate, claw linear;

wings oblong, 4–5(–5.4) × 1.5–2.1 mm;

keel obovate, (3.7–)4–5.3 × 2.4–3.7 mm;

stamens 5.2–6.8 mm;

filaments distinct to (1.5–)2–3 mm;

anthers (0.5–)0.6–0.8(–1) mm, connective gland-tipped.

Fruits

legumes, sessile, mostly tan, plump to compressed, obovoid, ovoid-ellipsoid, obovoid-ellipsoid, or obliquely obovoid or ellipsoid, indehiscent, often membranous proximally, thickened distally, gland-dotted, glabrous or pubescent.

Legumes

obliquely obovoid, 2.5 mm, with large glands distally, glabrous proximally, sparsely pilosulous distally.

Seeds

usually 1, rarely 2, chestnut to brownish or greenish, sometimes with brown or purple spots, somewhat compressed, oblong.

2–2.5 mm.

x

= 10.

Psorothamnus

Psorothamnus polydenius

Distribution
from USDA
sw United States; nw Mexico
[BONAP county map]
from USDA
sw United States
[WildflowerSearch map]
[BONAP county map]
Discussion

Species 9 (9 in the flora).

Psorothamnus and Psorodendron have equal priority. R. C. Barneby (1977d) was the first to accept Psorothamnus and cite Psorodendron as a synonym. Members of Psorothamnus often have been placed in Dalea. They differ from Dalea in the placement of the wing and keel petals, which are attached to the hypanthium rim and are not adnate to the stamen tube. R. C. Barneby (1977d) placed the nine species into four sections: Capnodendron Barneby (P. spinosus), Winnemucca Barneby (P. kingii), Xylodalea (S. Watson) Barneby (P. arborescens, P. fremontii, P. schottii), and Psorothamnus (P. emoryi, P. polydenius, P. scoparius, P. thompsoniae).

Asagraea Baillon 1870, not Lindley 1839, is an illegitimate name that pertains here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Bracteoles absent.
→ 2
2. Leaves unifoliolate or pinnate, leaflets 1 or 3; branches broomlike.
P. scoparius
2. Leaves mostly pinnate, leaflets 3–17(or 19), sometimes unifoliolate distally; branches divaricate.
→ 3
3. Leaves mostly pinnate, sometimes unifoliolate distally; terminal leaflets each longer than laterals; stems with antrorse-spreading hairs (when present).
P. emoryi
3. Leaves all pinnate; terminal leaflets each shorter than laterals; stems with retrorse hairs.
→ 4
4. Calyces with longer abaxial lobes; racemes dense to relatively loose, rachises.
0. 3–3 cm.
P. polydenius
4. Calyces with shorter abaxial lobes; racemes loose and open, rachises.
P. thompsoniae
1. Bracteoles present.
→ 5
5. Inflorescences with thornlike tips at anthesis.
→ 6
6. Trees (shrublike when young); anthers with gland-tipped connectives.
P. spinosus
6. Subshrubs (stems at intervals from creeping rootstocks); anthers without gland-tipped connectives.
P. kingii
5. Inflorescences without thornlike tips at anthesis.
→ 7
7. Leaves unifoliolate or pinnate; leaflet blades linear.
P. schottii
7. Leaves usually pinnate (leaflets 3–13), sometimes unifoliolate distally; leaflet blades often lanceolate, ovate, obovate, elliptic, rhombic-elliptic, rhombic-ovate, linear, linear-elliptic, or linear-oblanceolate.
→ 8
8. Legumes with scattered blister-glands (glands not confluent); leaflet blades silky-strigulose, villous-tomentulose to glabrate, loosely hairy, or glabrous.
P. arborescens
8. Legumes with numerous blister-glands (glands confluent into vertical ridges); leaflet blades strigulose, more strongly so adaxially.
P. fremontii
1. Calyx lobes 1.8–2.5 mm.
var. polydenius
1. Calyx lobes 3.6–5.2 mm.
var. jonesii
Source FNA vol. 11. Author: David M. Sutherland. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae Fabaceae > subfam. Faboideae > Psorothamnus
Sibling taxa
P. arborescens, P. emoryi, P. fremontii, P. kingii, P. schottii, P. scoparius, P. spinosus, P. thompsoniae
Subordinate taxa
P. arborescens, P. emoryi, P. fremontii, P. kingii, P. polydenius, P. schottii, P. scoparius, P. spinosus, P. thompsoniae
P. polydenius var. jonesii, P. polydenius var. polydenius
Synonyms Psorodendron Dalea polydenia
Name authority Rydberg in N. L. Britton et al.: N. Amer. Fl. 24: 45. (1919) (Torrey) Rydberg in N. L. Britton et al.: N. Amer. Fl. 24: 46. (1919) — (as polyadenius)
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