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dalea, indigo-bush

Mojave indigo-bush

Habit Shrubs, subshrubs, or trees, armed or unarmed, sterile shoots sometimes sharp-tipped. Shrubs.
Stems

erect, usually pubescent, rarely glabrate or glabrous, glandular nearly throughout, gland-dotted when young.

3–10 dm, gland-dotted distally when young, glands prickle-shaped, hairy to glabrate.

Branches

irregular, stiff;

sterile shoots often sharp-tipped.

Leaves

alternate, odd-pinnate or unifoliolate;

stipules present, caducous, usually triangular to subulate or linear, rarely obovate;

petiolate;

leaflets 1–17(or 19), stipels absent, blade margins entire or gland-crenulate, surfaces usually glandular-punctate abaxially, pubescent.

usually pinnate, sometimes unifoliolate distally, (0.3–)1–3.5(–5.5) cm;

leaflets (1 or 3)5–11, blades ovate, obovate, lanceolate, linear, elliptic, rhombic-elliptic, rhombic-ovate, or linear-oblanceolate, 2–14(–17) mm, terminal leaflet seldom longer than laterals, surfaces gland-dotted abaxially, silky-strigulose, villous-tomentulose to glabrate, loosely hairy, or glabrous.

Racemes

loose;

rachis without thornlike tip in anthesis, (1–)1.5–12 cm;

bracts subulate to linear, 2.5–4 mm;

bracteoles present.

Inflorescences

1–20+-flowered, terminal and axillary (sometimes leaf-opposed), usually racemes, rarely spikes;

bracts present, caducous;

bracteoles 0 or 2, at apex of pedicel or base of calyx.

Flowers

papilionaceous;

calyx campanulate, lobes 5, unequal;

ribs 10, not anastomosing into closed arches distally;

corolla usually blue, blue-purple, pink-purple, magenta-purple, violet-purple, violet, or bicolored, sometimes with yellow eye, rarely white;

banner blade differentiated from claw, reflexed less than 90°, claw shorter;

wings not adnate to keel, blades oblique basally, claws shorter, linear;

keel not strongly twisted, blades narrowly overlapping and adherent, oblique basally, blunt-tipped, claws linear;

stamens 10, monadelphous, equal or alternately short and long;

anthers dorsifixed;

style glabrous or pilosulous.

calyx 5–9 mm, pilosulous to minutely puberulent, strigulose, silky-canescent, or glabrate, tube 3–4.5 mm, ribs prominent, broader adaxial intervals with 2 rows, narrower one with 1 row of 3–5 glands, lobes broadly lanceolate, triangular, or lanceolate, abaxial lobe often shorter;

corolla indigo-blue to violet;

banner flabellate to flabellate-ovate, 6.3–10 mm, base cuneate to auriculate, claw cuneate;

wings oblong to oblong-ovate, 5.4–8.2 × 2.2–3.5 mm;

keel obliquely obovate, 5.8–8.4 × 3.3–4.9 mm;

stamens (5.5–)6.5–9.8 mm;

filaments distinct to (3–)3.5–5.6 mm;

anthers (0.8–)0.9–1.3 mm, connective not gland-tipped.

Fruits

legumes, sessile, mostly tan, plump to compressed, obovoid, ovoid-ellipsoid, obovoid-ellipsoid, or obliquely obovoid or ellipsoid, indehiscent, often membranous proximally, thickened distally, gland-dotted, glabrous or pubescent.

Legumes

ovoid-ellipsoid or obliquely ellipsoid, 8–10 mm, with scattered blister-glands, glands not confluent, glabrous or sparsely villosulous.

Seeds

usually 1, rarely 2, chestnut to brownish or greenish, sometimes with brown or purple spots, somewhat compressed, oblong.

(4.7–)5–6 mm.

x

= 10.

Psorothamnus

Psorothamnus arborescens

Distribution
from USDA
sw United States; nw Mexico
[BONAP county map]
from USDA
sw United States; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Species 9 (9 in the flora).

Psorothamnus and Psorodendron have equal priority. R. C. Barneby (1977d) was the first to accept Psorothamnus and cite Psorodendron as a synonym. Members of Psorothamnus often have been placed in Dalea. They differ from Dalea in the placement of the wing and keel petals, which are attached to the hypanthium rim and are not adnate to the stamen tube. R. C. Barneby (1977d) placed the nine species into four sections: Capnodendron Barneby (P. spinosus), Winnemucca Barneby (P. kingii), Xylodalea (S. Watson) Barneby (P. arborescens, P. fremontii, P. schottii), and Psorothamnus (P. emoryi, P. polydenius, P. scoparius, P. thompsoniae).

Asagraea Baillon 1870, not Lindley 1839, is an illegitimate name that pertains here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 4 (4 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Bracteoles absent.
→ 2
2. Leaves unifoliolate or pinnate, leaflets 1 or 3; branches broomlike.
P. scoparius
2. Leaves mostly pinnate, leaflets 3–17(or 19), sometimes unifoliolate distally; branches divaricate.
→ 3
3. Leaves mostly pinnate, sometimes unifoliolate distally; terminal leaflets each longer than laterals; stems with antrorse-spreading hairs (when present).
P. emoryi
3. Leaves all pinnate; terminal leaflets each shorter than laterals; stems with retrorse hairs.
→ 4
4. Calyces with longer abaxial lobes; racemes dense to relatively loose, rachises.
0. 3–3 cm.
P. polydenius
4. Calyces with shorter abaxial lobes; racemes loose and open, rachises.
P. thompsoniae
1. Bracteoles present.
→ 5
5. Inflorescences with thornlike tips at anthesis.
→ 6
6. Trees (shrublike when young); anthers with gland-tipped connectives.
P. spinosus
6. Subshrubs (stems at intervals from creeping rootstocks); anthers without gland-tipped connectives.
P. kingii
5. Inflorescences without thornlike tips at anthesis.
→ 7
7. Leaves unifoliolate or pinnate; leaflet blades linear.
P. schottii
7. Leaves usually pinnate (leaflets 3–13), sometimes unifoliolate distally; leaflet blades often lanceolate, ovate, obovate, elliptic, rhombic-elliptic, rhombic-ovate, linear, linear-elliptic, or linear-oblanceolate.
→ 8
8. Legumes with scattered blister-glands (glands not confluent); leaflet blades silky-strigulose, villous-tomentulose to glabrate, loosely hairy, or glabrous.
P. arborescens
8. Legumes with numerous blister-glands (glands confluent into vertical ridges); leaflet blades strigulose, more strongly so adaxially.
P. fremontii
1. Calyces 7–9 mm, lobes to 3–5.4 mm.
→ 2
2. Leaflets (3 or)5 or 7, blades ovate to obovate, elliptic, or broadly lanceolate.
var. arborescens
2. Leaflets (5 or)7–11, blades linear to linear-oblanceolate.
var. pubescens
1. Calyces 5–6.9 mm, lobes to 1.8–3.7 mm.
→ 3
3. Calyx tube glabrate or puberulent; leaflet blades lanceolate to rhombic-ovate or -elliptic, surfaces loosely hairy or glabrous.
var. minutifolius
3. Calyx tube silky-canescent externally; leaflet blades linear-oblanceolate to narrowly elliptic, surfaces silky-strigulose.
var. californicus
Source FNA vol. 11. Author: David M. Sutherland. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae Fabaceae > subfam. Faboideae > Psorothamnus
Sibling taxa
P. emoryi, P. fremontii, P. kingii, P. polydenius, P. schottii, P. scoparius, P. spinosus, P. thompsoniae
Subordinate taxa
P. arborescens, P. emoryi, P. fremontii, P. kingii, P. polydenius, P. schottii, P. scoparius, P. spinosus, P. thompsoniae
P. arborescens var. arborescens, P. arborescens var. californicus, P. arborescens var. minutifolius, P. arborescens var. pubescens
Synonyms Psorodendron Dalea arborescens
Name authority Rydberg in N. L. Britton et al.: N. Amer. Fl. 24: 45. (1919) (Torrey ex A. Gray) Barneby: Mem. New York Bot. Gard. 27: 33. (1977)
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