Psilostrophe villosa
Psilostrophe
woolly paperflower
paper-daisy, paperflower
arachno-villous (gray to gray-green).
erect to spreading, branched from bases or throughout (not scapiform).
basal and cauline or all cauline; alternate; petiolate or sessile;
blades spatulate to oblanceolate or linear, margins usually entire (sometimes toothed or lobed on larger rosette leaves), faces densely to sparsely arachno-villous or ± strigillose, often gland-dotted as well.
(0.5–)1–3(–5+) mm.
4–5(–7) mm.
cylindric to campanulate or obconic, 2–7 mm diam.
flat or convex, smooth or ± pitted (without setiform enations, sometimes gland-dotted), epaleate.
1–8, pistillate, fertile;
corollas yellow to orange (marcescent, spreading or reflexed in fruit).
5–8(–12).
5–25+, bisexual, fertile;
corollas yellow to orange, tubes shorter than narrowly cylindric throats, lobes 5, deltate (equal, papillate abaxially).
persistent, 5–12 in 1–2 series (erect in fruit, distinct, oblong to lanceolate, bases ± indurate).
in ± crowded, corymbiform arrays.
radiate, usually in compact, corymbiform arrays or glomerulate clusters (borne singly in P. cooperi).
usually glabrous, sometimes gland-dotted;
pappi of 4–6 lanceolate to lance-subulate scales 2.5–3+ mm.
cylindric to clavate or obpyramidal, sometimes weakly obcompressed (ray), all striate-ribbed, usually glabrous, sometimes gland-dotted (villous in P. gnaphalodes, sometimes hirtellous in P. tagetina);
pappi of 4–8 ± oblong or elliptic to lanceolate or lance-subulate, entire (lacerate in P. gnaphalodes) scales (without prominent midribs).
(2–)3(–5);
laminae 3–4(–6) mm, spreading in fruit.
= 16.
= 32.
Psilostrophe villosa
Psilostrophe
In trans-Pecos Texas, Psilostrophe villosa has been collected within 10 km or so of the Rio Grande; it probably occurs in Mexico. Within 100 km or so of the New Mexico-Texas border, P. villosa and P. tagetina intergrade; B. L. Turner et al. (1988) treated P. villosa as P. tagetina var. cerifera.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Species 7 (6 in the flora).
My treatment of psilostrophes closely follows that by R. C. Brown (1978). Psilostrophe mexicana R. C. Brown is known from Chihuahua and Durango.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Subshrubs or shrubs; stems pannose (white); heads borne singly; peduncles 35–60(–80+) mm | P. cooperi |
1. Biennials or perennials; stems arachno-villous (gray to gray-green) or strigillose (greenish); heads in corymbiform arrays; peduncles 0.5–25(–50) mm | → 2 |
2. Stems ± strigillose, (greenish, sometimes ± villous in proximal axils); ray laminaereflexed in fruit | P. sparsiflora |
2. Stems mostly arachno-villous (gray to gray-green); ray laminae spreading in fruit | → 3 |
3. Involucres 7–9(–12) mm; disc florets (10–)12–15(–20); pappus scales oblong to ovate, 1.5–2 mm | P. bakeri |
3. Involucres 4–6(–8) mm; disc florets 5–8(–12); pappus scales elliptic or lanceolate to linear-subulate, 2–3+ mm | → 4 |
4. Cypselae villous | P. gnaphalodes |
4. Cypselae usually glabrous, sometimes hirtellous and/or gland-dotted | → 5 |
5. Peduncles (3–)12–20(–40) mm; ray laminae (5–)7–14+ mm | P. tagetina |
5. Peduncles (0.5–)1–3(–5+) mm; ray laminae 3–4(–6) mm | P. villosa |
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