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sticky starwort, tuber starwort, tubered starwort

pseudostellaria, starwort, sticky starwort

Habit Herbs, perennial.
Taproots

absent, rhizomes usually with spherical or elongate, tuberous thickenings or vertical fleshy roots, rooting at nodes.

Rhizomes

with spherical or elongate tuberous thickenings 0.5–2.5 cm.

Stems

4-angled, 12–45(–60) cm, glabrous or stipitate-glandular throughout or at least in inflorescence, often densely so.

ascending to erect, simple or branched, terete or 4-angled.

Leaves

blades linear to linear-lanceolate or broadly lanceolate, (1.5–)2–10(–15) × 0.2–1.5(–2) cm, margins flat to briefly revolute, ± smooth or granular to serrulate, glabrous or stipitate-glandular.

connate proximally into sheath, sessile;

blade 1-veined, linear to lanceolate or elliptic, not succulent, apex acute.

Inflorescences

open cymes, flowers often proliferating with age.

terminal, open cymes, or flowers solitary;

bracts paired, smaller, herbaceous or scarious.

Pedicels

recurved to reflexed from base in fruit, uniformly stipitate-glandular.

recurved to reflexed from base or abruptly bent downward near distal end in fruit.

Flowers

sepals lanceolate to narrowly ovate, 3–5.5(–7) × 0.8–2 mm, stipitate-glandular, often densely so;

petals 7–9.5 × 3–4 mm, apex notch 1–2 mm deep, lobes broadly rounded;

anthers 10, purple;

styles 3.5–4.5 mm;

stigmas terminal, 0.1–0.2 mm.

perianth and androecium hypogynous;

sepals 5, distinct, green, narrowly ovate to lanceolate, 3–7 mm, herbaceous, margins white, scarious, apex obtuse or acute to acuminate, not hooded;

petals 5 [absent in cleistogamous flowers], white, not clawed, blade apex 2-fid to V-shaped notch 1/10–1/5 of length;

nectaries adnate to base of filaments opposite sepals, circular, thickened, 2–21/2 times filament width;

stamens 5 or 10, arising from base of ovary;

filaments distinct;

styles 3, capitate to clavate, 2–4.5 mm, glabrous proximally;

stigmas 3, terminal or linear along adaxial surface of styles, minutely papillate (30x).

Capsules

4.5–5 mm.

ovoid, opening by 6, ± 2–3 times recoiled valves;

carpophore absent.

Seeds

1–3, reddish brown, broadly elliptic, ± plump, 2–3.4 mm;

tubercles conic to elongate, rounded.

1–3, red-brown or brown, circular to oblong or elliptic, plump or laterally compressed, tuberculate, marginal wing absent, appendage absent.

x

= 8.

2n

= 96.

Pseudostellaria jamesiana

Pseudostellaria

Phenology Flowering summer.
Habitat Meadows, sagebrush-grasslands, dry understory of aspen and coniferous forests
Elevation 600-3400 m. (2000-11200 ft.)
Distribution
from FNA
AZ; CA; CO; ID; MT; NM; NV; OR; TX; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
from USDA
w United States; se Europe; e Asia
[BONAP county map]
Discussion

Species 21 (3 in the flora).

We follow W. A. Weber and R. L. Hartman (1979) in recognizing Pseudostellaria in North America. The combination of a V-shaped petal sinus cut to only 1/10–1/5 of the petal length, often tuberous rhizomes and thickened roots, and the more or less recoiled capsule valves segregates our species from Stellaria. The relationship between the North American and Asian species requires further investigation. While the seeds of P. sierrae and P. oxyphylla closely resemble those of P. rupestris (Turczaninow) Pax, the type species, the presence of five (rather than ten) stamens, absence of tubers, and absence of cleistogamous flowers complicate placing our species in the current infrageneric classification (M. Mizushina 1965).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Pedicels and sepals uniformly stipitate-glandular, often densely so; anthers 10, purple
P. jamesiana
1. Pedicels glabrous or with narrow internodal line of hairs; sepals glabrous or with margins ciliolate in proximal 2; anthers 5, yellow
→ 2
2. Stems and pedicels each with narrow internodal line of hairs; leaf blades 6-12 cm; inflorescences with flowers paired (single by abortion), pedicellate, in distal 3-7 axils or in terminal cymes
P. oxyphylla
2. Stems and pedicels glabrous; leaf blades 0.7-3.5 cm; inflorescences with flowers solitary, terminal, sometimes on 1-2 axillary branches
P. sierrae
Source FNA vol. 5, p. 115. FNA vol. 5, p. 114. Authors: Ronald L. Hartman, Richard K. Rabeler.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Pseudostellaria Caryophyllaceae > subfam. Alsinoideae
Sibling taxa
P. oxyphylla, P. sierrae
Subordinate taxa
P. jamesiana, P. oxyphylla, P. sierrae
Synonyms Stellaria jamesiana, Alsine glutinosa, Arenaria jamesiana Krascheninikovia
Name authority (Torrey) W. A. Weber & R. L. Hartman: Phytologia 44: 314. (1979) Pax: in H. G. A. Engler and K. Prantl, Nat. Pflanzenfam. ed. 2, 16c: 318. (1934)
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