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pseudoleskeella moss

pseudoleskeella moss

Habit Plants dark olive green or rarely yellow-green. Plants small, in thin mats or patches, yellow- to red-green, red-brown, green, or olive green.
Stem(s)

leaves ± catenulate when dry, not homomallous, ovate, abruptly narrowed to apex, concave, not plicate, 0.4–1.2 mm, 1.5–2:1;

base flaring;

margins plane or rarely recurved proximally, entire;

apex short-acuminate, sometimes falcate;

costa usually double, or single, 2-fid, or sometimes ecostate, weak, not reaching mid leaf;

medial laminal cells 1–2(–3):1, walls firm, not incrassate.

and branch leaves differentiated.;

stem leaves catenulate or not, appressed to erect when dry, erect-spreading when moist, ovate to lanceolate, not to sometimes 2-plicate;

margins plane or recurved, entire to serrulate distally;

apex abruptly to gradually acuminate, hair-point absent;

costa single, double, 2-fid, or rarely ecostate, strong to weak, short to percurrent, not sinuate, sometimes somewhat obscure distally;

alar cells not well differentiated, quadrate to short-rectangular;

proximal laminal cells 1–2:1, walls not or obscurely pitted;

medial cells isodiametric, 2–3:1, smooth, walls firm to incrassate.

Branch leaves

ovate to suborbicular, 0.4–0.8 mm;

laminal cell walls moderately incrassate.

more ovate, smaller;

apex often especially recurved or falcate;

costa usually weaker;

medial laminal cells usually shorter.

Seta

red-brown, 0.1–0.2 cm.

0.1–0.3 cm.

Sexual condition

dioicous;

perichaetial leaves pale translucent, appressed to rarely recurved, longer, apex more acuminate.

Capsule

inclined, subcylindric, asymmetric, 1.5–2.5(–3) mm.

erect or inclined, subcylindric to cylindric, curved-asymmetric or sometimes symmetric;

annulus present;

operculum conic, short-rostrate;

peristome well developed;

exostome teeth lanceolate or linear-lanceolate, finely horizontally striate;

endostome basal membrane moderately high, segments slender, lanceolate to filiform, cilia present, usually well developed.

Spores

10–18 µm.

8–18 µm, finely papillose.

Stoloniferous

leaves broadly ovate or suborbicular.

Specialized

asexual reproduction absent.

Pseudoleskeella tectorum

Pseudoleskeella

Phenology Capsules mature summer.
Habitat Shaded calcareous rock, shaded tree bases or wood
Elevation low to high elevations (0-4000 m) (low to high elevations (0-13100 ft))
Distribution
from FNA
AK; AZ; CA; CO; ID; KS; MI; MN; MT; ND; NE; NM; NV; OR; SD; UT; WY; AB; BC; NL; NT; NU; ON; QC; SK; Greenland; Eurasia
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North America; Mexico; Eurasia
Discussion

Pseudoleskeella tectorum is distinguished by the dark green, tightly appressed, catenulate leaves, and weak costa. Thin mats of this species can be become quite extensive over time, covering the faces and overhangs of shaded boulders and cliffs.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 6 (4 in the flora).

Pseudoleskeella is found mostly in cool-temperate, montane, arctic-alpine, and boreal-temperate regions; the plants are small, slender, and appressed, with many branches, smooth laminal cells, no paraphyllia, straight to flexuose setae, and a relatively weak costa, often 2-fid. The older stoloniferous leaves are distant, often denuded, scalelike, and sometimes hyaline; occasionally some distal laminal cells become weakly prorate. However, the species seem to be quite different from each other morphologically beyond these few shared traits. Molecular studies have shown that P. serpentinensis is not related to the other species; rather it appears to be closest to Heterocladium (Pterigynandraceae) (M. S. Ignatov et al. 2007). Following P. S. Wilson and D. H. Norris (1989), P. catenulata (Bridel ex Schrader) Kindberg and P. papillosa (Lindberg) Kindberg are excluded from North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stem leaves ± catenulate when dry; costae usually double, not reaching mid leaf; stems tightly appressed to substrate.
P. tectorum
1. Stem leaves not catenulate when dry; costae usually single, rarely 2-fid, usually to mid leaf or sometimes percurrent; stems not tightly appressed to substrate
→ 2
2. Plants green to yellow-green; leaves distinctly plicate; bases flaring, cordate; margins recurved to mid leaf or beyond, distinctly serrulate distally.
P. arizonae
2. Plants green to red; leaves not or weakly plicate; bases not flaring or cordate; margins plane throughout or recurved proximally, ± entire or finely serrulate distally
→ 3
3. Stem leaves broadly lanceolate to ovate-lanceolate, not homomallous, gradually narrowed to apex; apices not falcate; costae usually beyond mid leaf, rarely 2-fid; plants green, red-green, or orange-green; branch leaf laminal cell walls thin.
P. rupestris
3. Stem leaves ovate to lanceolate, homomallous, abruptly narrowed to apex; apices sometimes falcate; costae short and 2-fid to single and reaching mid leaf or sometimes beyond; plants red, red-brown, or green; branch leaf laminal cell walls incrassate.
P. serpentinensis
Source FNA vol. 28, p. 365. FNA vol. 28, p. 361. Author: John R. Spence.
Parent taxa Leskeaceae > Pseudoleskeella Leskeaceae
Sibling taxa
P. arizonae, P. rupestris, P. serpentinensis
Subordinate taxa
P. arizonae, P. rupestris, P. serpentinensis, P. tectorum
Synonyms Hypnum tectorum, Pseudoleskea tectorum
Name authority (Funck ex Bridel) Kindberg ex Brotherus: in H. G. A. Engler and K. Prantl, Nat. Pflanze n fam. 229[I,3]: 997. (1907) Kindberg: Eur. N. Amer. Bryin., 47. (1897)
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