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pseudoleskea moss

pseudoleskea moss

Habit Plants medium-sized to large, in thick mats, green, yellow-green, or orange-green. Plants small, in thin open to thick mats, green or yellow-green.
Stems

with branches robust, julaceous, apices usually curving up;

central strand present;

paraphyllia many, filamentous to foliose, branched.

with branches slender, not or rarely julaceous, apices straight to rarely curving up;

central strand present;

paraphyllia many, filamentous to foliose, not branched.

Leaves

appressed to somewhat erect when dry, erect-spreading when moist, glossy or dull, ovate to ovate-lanceolate, asymmetric, usually falcate to falcate-secund, 0.6–2(–2.4) mm;

margins narrowly recurved to mid leaf or to near acumen;

apex abruptly acute to short- or long-acuminate, hair-point absent;

costa subpercurrent to percurrent, yellow-green, sometimes sinuate;

alar cells transversely elongate to quadrate, region small to medium-sized;

medial laminal cells homogeneous, short-rhomboidal, elliptic, or fusiform, to 40 µm, 2–3(–4):1, pellucid to opaque, prorate to near base, lumina larger than 10 µm, walls thin or rarely firm, not pitted;

apical cells 2–3:1;

juxtacostal cells somewhat shorter than more distal cells, walls not pitted.

appressed to somewhat erect when dry, erect-spreading when moist, glossy, ovate-lanceolate to lanceolate, symmetric, not or rarely falcate except at apices, 0.9–2 mm;

margins recurved to just before acumen;

apex gradually and slenderly long-acuminate, hair-point absent;

costa percurrent, green to yellow-green, not sinuate;

alar cells transversely elongate to quadrate, region small;

medial laminal cells elongate-rhomboidal to fusiform or somewhat vermicular, to 30 µm, 2–3(–4):1, pellucid, strongly papillose over lumen, papillae usually off-centered to distal edge, rarely some cells prorate, walls firm to incrassate, not pitted;

juxtacostal cells somewhat shorter than more distal cells, walls not pitted.

Capsule

erect to suberect, symmetric, 1–2 mm;

endostome basal membrane 1/4–1/3 exostome length, segments shorter than exostome, cilia usually present, 1 or 2, sometimes reduced.

erect, symmetric, 1–3 mm;

endostome basal membrane 1/4 exostome length, segments shorter than exostome, cilia absent or rudimentary.

Spores

16–22 µm.

16–20 µm.

Pseudoleskea radicosa

Pseudoleskea stenophylla

Phenology Capsules mature spring–summer (May–Aug).
Habitat Twigs and branches of shrubs or small trees, tree bases of Acer or Alnus, along streams
Elevation moderate to high elevations (500-2100 m) (moderate to high elevations (1600-6900 ft))
Distribution
North America; Eurasia; Atlantic Islands (Iceland)
[WildflowerSearch map]
from FNA
AK; CA; ID; MT; OR; UT; WA; AB; BC; NL; NS; QC; w Eurasia
Discussion

Varieties 3 (3 in the flora).

Pseudoleskea radicosa is a common and variable species found at moderate to high elevations; the capsules mature in summer. Two of the three varieties are quite distinctive. Pseudoleskea radicosa is closely related to P. incurvata but differs in the longer, more homogeneous, thin-walled, and wider laminal cells. Combined ecological, molecular and morphological studies of the varieties are needed to determine their distinctiveness.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pseudoleskea stenophylla is distinguished by small plants growing on bark and twigs of shrubs and trees, leaves with a long, slender acumen half the leaf length, and elongate laminal cells with papillae over the lumen but usually off-centered.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants large; stem apices hooked; branches few or absent; leaves 1.6-2.3 mm, strongly falcate-secund, pluriplicate; apices long-acuminate.
var. denudata
1. Plants small to medium-sized; stem apices not distinctly hooked; branches many; leaves usually less than 1.6 mm, straight, falcate, or falcate-secund, 2-plicate; apices predominantly short-acuminate to acute
→ 2
2. Plants medium-sized; leaves weakly to moderately concave, not incurved, 1.2-1.7 mm; apices usually long-acuminate; medial laminal cells opaque.
var. radicosa
2. Plants small; leaves strongly concave, often incurved, 0.8-1.3 mm; apices acute to short-acuminate; medial laminal cells pellucid.
var. compacta
Source FNA vol. 28, p. 359. FNA vol. 28, p. 361.
Parent taxa Leskeaceae > Pseudoleskea Leskeaceae > Pseudoleskea
Sibling taxa
P. atricha, P. baileyi, P. incurvata, P. patens, P. saviana, P. stenophylla, P. tribulosa
P. atricha, P. baileyi, P. incurvata, P. patens, P. radicosa, P. saviana, P. tribulosa
Subordinate taxa
P. radicosa var. compacta, P. radicosa var. denudata, P. radicosa var. radicosa
Synonyms Hypnum radicosum, Lescuraea radicosa Lescuraea stenophylla
Name authority (Mitten) Macoun & Kindberg: Cat. Canad. Pl., Musci, 181. (1892) Renauld & Cardot: Bot. Centra l bl. 44: 421. (1890)
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