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pseudoleskea moss

patent leskea, patent mountain leskea, pseudoleskea moss

Habit Plants medium-sized to large, in thick mats, green, yellow-green, or orange-green. Plants small to medium-sized, in thin mats, dark green, olive green, black-green, or rarely yellow-green.
Stems

with branches robust, julaceous, apices usually curving up;

central strand present;

paraphyllia many, filamentous to foliose, branched.

with branches slender to robust, julaceous, apices sometimes curving up;

central strand present;

paraphyllia many, filamentous to foliose, not branched.

Leaves

appressed to somewhat erect when dry, erect-spreading when moist, glossy or dull, ovate to ovate-lanceolate, asymmetric, usually falcate to falcate-secund, 0.6–2(–2.4) mm;

margins narrowly recurved to mid leaf or to near acumen;

apex abruptly acute to short- or long-acuminate, hair-point absent;

costa subpercurrent to percurrent, yellow-green, sometimes sinuate;

alar cells transversely elongate to quadrate, region small to medium-sized;

medial laminal cells homogeneous, short-rhomboidal, elliptic, or fusiform, to 40 µm, 2–3(–4):1, pellucid to opaque, prorate to near base, lumina larger than 10 µm, walls thin or rarely firm, not pitted;

apical cells 2–3:1;

juxtacostal cells somewhat shorter than more distal cells, walls not pitted.

appressed to julaceous when dry, erect-spreading to patent when moist, dull, ovate, asymmetric, not or rarely falcate, (0.6–)0.7–1 mm;

margins recurved proximally;

apex abruptly acute to rarely short-acuminate, hair-point absent;

costa not reaching acumen, green to orange-green, somewhat sinuate, terminal abaxial cells not or weakly projecting, not abaxially strongly dentate;

alar cells quadrate to transversely elongate, region large;

medial laminal cells quadrate, to 18 µm, 1(–2):1, opaque, moderately to strongly low rounded papillose over lumen or papillae sometimes off-centered, walls incrassate, not pitted;

proximal juxtacostal cells usually shorter than medial, walls pitted or not.

Capsule

erect to suberect, symmetric, 1–2 mm;

endostome basal membrane 1/4–1/3 exostome length, segments shorter than exostome, cilia usually present, 1 or 2, sometimes reduced.

inclined, asymmetric, 0.5–2 mm;

endostome basal membrane 1/3–1/2 exostome length, segments shorter than exostome, cilia well developed.

Spores

16–22 µm.

10–14 µm.

Pseudoleskea radicosa

Pseudoleskea patens

Phenology Capsules mature summer (Jul–Aug).
Habitat Shaded to exposed rock and outcrops, mineral soil, near streams
Elevation moderate to high elevations (900-3500 m) (moderate to high elevations (3000-11500 ft))
Distribution
North America; Eurasia; Atlantic Islands (Iceland)
[WildflowerSearch map]
from FNA
AK; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC; NL; NS; NT; Eurasia
Discussion

Varieties 3 (3 in the flora).

Pseudoleskea radicosa is a common and variable species found at moderate to high elevations; the capsules mature in summer. Two of the three varieties are quite distinctive. Pseudoleskea radicosa is closely related to P. incurvata but differs in the longer, more homogeneous, thin-walled, and wider laminal cells. Combined ecological, molecular and morphological studies of the varieties are needed to determine their distinctiveness.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pseudoleskea patens is relatively small and usually dull, dark, or olive green, with short laminal cells with rounded centrally located papillae; it differs from P. tribulosa by the recurved margins, weaker and lower laminal cell papillae lacking projecting apical salient, larger leaves with stronger plicae, and lack of the Morningstar-shaped projections of the abaxial apex cells of the costa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants large; stem apices hooked; branches few or absent; leaves 1.6-2.3 mm, strongly falcate-secund, pluriplicate; apices long-acuminate.
var. denudata
1. Plants small to medium-sized; stem apices not distinctly hooked; branches many; leaves usually less than 1.6 mm, straight, falcate, or falcate-secund, 2-plicate; apices predominantly short-acuminate to acute
→ 2
2. Plants medium-sized; leaves weakly to moderately concave, not incurved, 1.2-1.7 mm; apices usually long-acuminate; medial laminal cells opaque.
var. radicosa
2. Plants small; leaves strongly concave, often incurved, 0.8-1.3 mm; apices acute to short-acuminate; medial laminal cells pellucid.
var. compacta
Source FNA vol. 28, p. 359. FNA vol. 28, p. 359.
Parent taxa Leskeaceae > Pseudoleskea Leskeaceae > Pseudoleskea
Sibling taxa
P. atricha, P. baileyi, P. incurvata, P. patens, P. saviana, P. stenophylla, P. tribulosa
P. atricha, P. baileyi, P. incurvata, P. radicosa, P. saviana, P. stenophylla, P. tribulosa
Subordinate taxa
P. radicosa var. compacta, P. radicosa var. denudata, P. radicosa var. radicosa
Synonyms Hypnum radicosum, Lescuraea radicosa Lescuraea patens
Name authority (Mitten) Macoun & Kindberg: Cat. Canad. Pl., Musci, 181. (1892) (Lindberg) Kindberg: Canad. Rec. Sci. 6: 20. (1894)
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