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pseudoleskea moss

pseudoleskea moss

Habit Plants medium-sized to large, in thick mats, green, yellow-green, orange-green, or rarely black-green. Plants large, in thick, stiff mats, green, yellow-green, orange-green, or black-green.
Stems

with branches robust, julaceous, apices curving up;

central strand present;

paraphyllia many, filamentous to foliose, branched.

with branches robust, julaceous, apices curving up;

central strand present;

paraphyllia many, filamentous, not branched.

Leaves

appressed to somewhat erect when dry, erect-spreading when moist, glossy or dull, ovate to ovate-lanceolate, asymmetric, usually falcate to falcate-secund, 0.8–2(–2.5) mm;

margins strongly and narrowly recurved to near acumen;

apex abruptly acute to short- or rarely long-acuminate, hair-point absent;

costa subpercurrent to percurrent, orange-green, sometimes sinuate;

alar cells transversely elongate to quadrate, region large;

medial laminal cells heterogeneous, elliptic or isodiametric, to 25 µm, 1–2(–3):1, opaque or sometimes pellucid, prorate to near base, lumina smaller than 8 µm, walls firm to strongly incrassate, not pitted;

apical cells 1–2:1;

juxtacostal cells somewhat shorter than more distal cells, walls not pitted.

appressed to julaceous when dry, erect-spreading when moist, dull, ovate, asymmetric, rarely somewhat falcate, (0.5–)1–1.2 mm;

margins recurved proximally;

apex gradually acute to rarely short-acuminate, hair-point absent;

costa percurrent, green to yellow-green, somewhat sinuate;

alar cells transversely elongate, quadrate, or short-rectangular, region medium-sized;

medial laminal cells elongate-rhomboidal, to 40 µm, 3–4:1, opaque, distinctly prorate, walls incrassate, strongly pitted;

juxtacostal cells usually shorter than medial cells, walls pitted or not.

Capsule

erect to suberect, symmetric, 1–2.5 mm;

endostome basal membrane 1/4–1/3 exostome length, segments shorter than exostome, cilia 1 or 2.

inclined to suberect, asymmetric, 0.5–1.2 mm;

endostome basal membrane 1/3–1/2 exostome length, segments shorter than exostome, cilia well developed to occasionally rudimentary.

Spores

10–20 µm.

(10–)12–18(–20) µm.

Pseudoleskea incurvata

Pseudoleskea atricha

Phenology Capsules mature summer (Jul–Aug).
Habitat Subalpine and alpine rock outcrops, mineral soil
Elevation moderate to high elevations (600-2300 m) (moderate to high elevations (2000-7500 ft))
Distribution
North America; Eurasia; North Temperate Zone; moderate to high elevations
[WildflowerSearch map]
from FNA
AK; WA; BC
Discussion

Varieties 11 (3 in the flora).

Pseudoleskea incurvata is a complex and highly variable species; the capsules mature in summer. Most plants can be distinguished from P. radicosa by the thick-walled proximal and medial laminal cells that are mostly short, and by the relatively short cells in the acumen. Although there are intermediate specimens, the three varieties are often quite distinct. Combined ecological, molecular and morphological studies of the varieties are needed to determine their distinctiveness.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pseudoleskea atricha is a local high elevation endemic species characterized by ovate leaves with an acute but not acuminate apex, and incrassate laminal cells that are strongly pitted. Only P. baileyi also has strongly pitted cells, but it differs in having leaves with long acumina and hair-points, and stems lacking paraphyllia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants large; leaves 1.5-2.5 mm, distinctly falcate-secund; apices long-acuminate; laminal cell prorulae indistinct.
var. gigantea
1. Plants small to medium-sized; leaves smaller than1.5 mm, straight, falcate, or falcate-secund; apices usually short-acuminate to acute; laminal cell prorulae distinct
→ 2
2. Leaves 1-1.5 mm; apices acute to short-acuminate to acuminate; medial laminal cells elliptic, corners rounded.
var. incurvata
2. Leaves 0.8-1.1 mm; apices abruptly acute to short-acuminate; medial laminal cells isodiametric, corners angular.
var. tenuiretis
Source FNA vol. 28, p. 357. FNA vol. 28, p. 356.
Parent taxa Leskeaceae > Pseudoleskea Leskeaceae > Pseudoleskea
Sibling taxa
P. atricha, P. baileyi, P. patens, P. radicosa, P. saviana, P. stenophylla, P. tribulosa
P. baileyi, P. incurvata, P. patens, P. radicosa, P. saviana, P. stenophylla, P. tribulosa
Subordinate taxa
P. incurvata var. gigantea, P. incurvata var. incurvata, P. incurvata var. tenuiretis
Synonyms Leskea incurvata, Lescuraea incurvata P. atrovirens var. atricha, Lescuraea atricha
Name authority (Hedwig) Loeske: Hedwigia 50: 313. (1911) (Kindberg) Kindberg: Ottawa Naturalist 7: 20. (1893)
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