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Heller's cudweed, Heller's rabbit-tobacco

cliff cudweed

Habit Annuals (fragrant), 30–100 cm; fibrous-rooted (roots relatively thick and lignescent). Annuals, 4–15(–30) cm; taprooted.
Stems

greenish, glandular-villous (without persistent tomentum, stipitate glands mostly 0.3–1 mm, often variable, stalks broadened toward bases, about equaling gland widths).

(filiform) persistently tomentose (indument a loose, envelope-like, transparent haze of extremely thin hairs, doubling apparent stem width), not glandular.

Leaf

blades mostly oblong-lanceolate, 2.5–7 cm × 4–20 mm, bases not clasping, not decurrent, margins flat, faces bicolor, abaxial white to gray with lightly persistent tomentum, adaxial green, both minutely stipitate-glandular.

blades elliptic-oblanceolate to oblanceolate, 0.5–3 cm × 2–6 mm (largest at midstem), bases not clasping, not decurrent, margins flat, faces concolor, green, thinly arachnoid-tomentose to glabrate, not glandular (veiny reticulum evident).

Involucres

campanulate, 6–7 mm.

turbinate, 4–5 mm.

Pistillate florets

83–107.

25–28.

Bisexual florets

9–15.

6–7.

Phyllaries

in 4–6 series, white (opaque, shiny), ovate to ovate-oblong or oblong, ± tomentose.

in 3(–4) series, whitish to slightly tawny (hyaline, shiny), narrowly triangular to narrowly oblong-triangular, glabrous.

Heads

in corymbiform arrays.

(2–4) in terminal, capitate clusters (usually immediately subtended by distalmost cauline leaf, clusters sometimes in subcorymbiform arrays).

Cypselae

ridged, smooth.

not ridged, smooth.

Pseudognaphalium helleri

Pseudognaphalium saxicola

Phenology Flowering Sep–Oct(–Nov). Flowering (Jul–)Aug–Sep.
Habitat Dry woods and openings, clay and sandy clay, sand hills Mostly bare sandstone cliff faces, ledges, and cracks, s- to e-facing, commonly shaded
Elevation 10–300 m (0–1000 ft) 200–300 m (700–1000 ft)
Distribution
from FNA
AL; AR; FL; GA; LA; MS; NC; OK; SC; TN; TX; VA
[WildflowerSearch map]
[BONAP county map]
from FNA
WI
Discussion

Pseudognaphalium helleri and P. micradenium are similar to P. obtusifolium in most features; both differ in their glandular stems without the persistent whitish tomentum of P. obtusifolium.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pseudognaphalium saxicola probably is an evolutionary derivative of P. obtusifolium. Plants of P. saxicola are relatively small and have relatively few, relatively small heads and occur in a specialized habitat; they constitute the only narrowly endemic species of Pseudognaphalium in the United States. Depauperate individuals of P. obtusifolium from localities over its whole geographic range may sometimes be as short as 5–10 cm and similar in habit to P. saxicola; such plants differ from P. saxicola in their close and denser stem vestiture, bicolor and relatively narrow leaves, larger heads with greater numbers of pistillate florets, and broader phyllaries with rounded apices.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 19, p. 421. FNA vol. 19, p. 420.
Parent taxa Asteraceae > tribe Gnaphalieae > Pseudognaphalium Asteraceae > tribe Gnaphalieae > Pseudognaphalium
Sibling taxa
P. arizonicum, P. austrotexanum, P. beneolens, P. biolettii, P. californicum, P. canescens, P. jaliscense, P. leucocephalum, P. luteoalbum, P. macounii, P. micradenium, P. microcephalum, P. obtusifolium, P. pringlei, P. ramosissimum, P. roseum, P. saxicola, P. stramineum, P. thermale, P. viscosum
P. arizonicum, P. austrotexanum, P. beneolens, P. biolettii, P. californicum, P. canescens, P. helleri, P. jaliscense, P. leucocephalum, P. luteoalbum, P. macounii, P. micradenium, P. microcephalum, P. obtusifolium, P. pringlei, P. ramosissimum, P. roseum, P. stramineum, P. thermale, P. viscosum
Synonyms Gnaphalium helleri, Gnaphalium obtusifolium var. helleri Gnaphalium saxicola, Gnaphalium obtusifolium var. saxicola, P. obtusifolium var. saxicola
Name authority (Britton) Anderberg: Opera Bot. 104: 147. (1991) (Fassett) H. E. Ballard & Feller: Sida 21: 777. (2004)
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