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Heller's cudweed, Heller's rabbit-tobacco

Bioletti's rabbit-tobacco, two-color rabbit-tobacco, two-tone everlasting

Habit Annuals (fragrant), 30–100 cm; fibrous-rooted (roots relatively thick and lignescent). Perennials (fragrant), 20–70(–120) cm; taprooted.
Stems

greenish, glandular-villous (without persistent tomentum, stipitate glands mostly 0.3–1 mm, often variable, stalks broadened toward bases, about equaling gland widths).

(sometimes lignescent near bases) proximally glabrescent, distally persistently tomentose, at least in the region of the heads, not glandular.

Leaf

blades mostly oblong-lanceolate, 2.5–7 cm × 4–20 mm, bases not clasping, not decurrent, margins flat, faces bicolor, abaxial white to gray with lightly persistent tomentum, adaxial green, both minutely stipitate-glandular.

blades (not crowded, internodes mostly 5+ mm), oblong-oblanceolate to oblanceolate (the distal lanceolate), 1.5–5(–8) cm × 4–10(–15) mm, bases auriculate-clasping, not decurrent, margins flat or slightly revolute, often undulate, faces bicolor (at least basal and proximal cauline), abaxial white-tomentose, adaxial bright green and ± densely glandular.

Involucres

campanulate, 6–7 mm.

turbinate-campanulate, 5–5.5(–6) mm.

Pistillate florets

83–107.

41–73.

Bisexual florets

9–15.

5–13.

Phyllaries

in 4–6 series, white (opaque, shiny), ovate to ovate-oblong or oblong, ± tomentose.

in 4–5 series, white or sometimes slightly pinkish (opaque, shiny), ovate to oblong-ovate or oblong (often longitudinally wrinkled or grooved), glabrous.

Heads

in corymbiform arrays.

in corymbiform arrays.

Cypselae

ridged, smooth.

ridged, smooth.

Pseudognaphalium helleri

Pseudognaphalium biolettii

Phenology Flowering Sep–Oct(–Nov). Flowering Apr–Jun(–Oct).
Habitat Dry woods and openings, clay and sandy clay, sand hills Rocky slopes, roadsides, sandy plains with Larrea, coastal strand, matorral, and chaparral
Elevation 10–300 m (0–1000 ft) 5–600(–1200) m (0–2000(–3900) ft)
Distribution
from FNA
AL; AR; FL; GA; LA; MS; NC; OK; SC; TN; TX; VA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico (Baja California, Baja California Sur)
[WildflowerSearch map]
[BONAP county map]
Discussion

Pseudognaphalium helleri and P. micradenium are similar to P. obtusifolium in most features; both differ in their glandular stems without the persistent whitish tomentum of P. obtusifolium.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 19, p. 421. FNA vol. 19, p. 423.
Parent taxa Asteraceae > tribe Gnaphalieae > Pseudognaphalium Asteraceae > tribe Gnaphalieae > Pseudognaphalium
Sibling taxa
P. arizonicum, P. austrotexanum, P. beneolens, P. biolettii, P. californicum, P. canescens, P. jaliscense, P. leucocephalum, P. luteoalbum, P. macounii, P. micradenium, P. microcephalum, P. obtusifolium, P. pringlei, P. ramosissimum, P. roseum, P. saxicola, P. stramineum, P. thermale, P. viscosum
P. arizonicum, P. austrotexanum, P. beneolens, P. californicum, P. canescens, P. helleri, P. jaliscense, P. leucocephalum, P. luteoalbum, P. macounii, P. micradenium, P. microcephalum, P. obtusifolium, P. pringlei, P. ramosissimum, P. roseum, P. saxicola, P. stramineum, P. thermale, P. viscosum
Synonyms Gnaphalium helleri, Gnaphalium obtusifolium var. helleri Gnaphalium bicolor
Name authority (Britton) Anderberg: Opera Bot. 104: 147. (1991) Anderberg: Opera Bot. 104: 147. (1991)
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