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Kern cinquefoil, Wheeler's cinquefoil

potentille de Vahl, Vahl's cinquefoil

Habit Plants densely tufted to cushion-forming.
Caudex branches

usually stout, columnar, sheathed with marcescent whole leaves.

Stems

0.2–2.5 dm.

erect, (0.2–)0.3–0.8(–1) dm, lengths 2–4(–5) times basal leaves.

Basal leaves

palmate, 1.5–9.5 cm;

petiole 1–7 cm, long hairs abundant to dense, spreading to appressed, 1–2 mm, weak to stiff, glands ± abundant;

leaflets 5, central cuneate-elliptic to obovate, 0.5–2.5 × 0.3–1.5 cm, scarcely to distinctly petiolulate, distal 1/3 of margins evenly incised ± 1/4 to midvein, teeth 2–4(–5) per side, surfaces gray-green, long hairs abundant to dense, 1–1.5 mm, glands ± abundant, sometimes obscured.

1–3 cm;

petiole 0.5–2 cm, long hairs common to abundant, ascending to spreading, 0.5–1.5(–2) mm, soft to weak, smooth, short hairs absent, crisped/short-cottony hairs sparse to abundant, glands sparse to common;

leaflets separate to slightly overlapping, central obovate, 0.8–1.2(–1.5) × 0.5–1(–1.5) cm, sessile or subsessile, base cuneate, margins revolute, distal ± 1/2 incised (1/3–)1/2–3/4 to midvein, teeth 2–3 per side, ± approximate to distant, surfaces ± dissimilar, abaxial yellowish white or grayish, long hairs 1–2 mm, cottony-crisped hairs ± dense, adaxial green to yellowish gray, long(/short) hairs common to abundant, crisped hairs absent or sparse.

Cauline leaves

0–1(–2).

Inflorescences

1–20-flowered.

1–2(–3)-flowered.

Pedicels

0.5–1.5 cm.

0.5–1.5 cm in flower, to 3.5 cm in fruit.

Flowers

epicalyx bractlets lanceolate to ovate-elliptic, 1–3.5 × 1–1.5 mm;

hypanthium 2–4 mm diam.;

sepals 2–5 mm, apex ± acute;

petals ± paler abaxially, bright yellow adaxially, ± obcordate, 3–6 × 2–5 mm;

filaments 1–2 mm, anthers 0.5 mm;

carpels ca. 20, styles (1.2–)1.5–2 mm.

epicalyx bractlets broadly ovate, 2.5–4(–5) × 1.5–3 mm, ± as wide as sepals, margins flat or revolute, red glands absent;

hypanthium 2–4 mm diam.;

sepals 2.5–5(–6) mm, apex subacute to acute;

petals 8–10 × 7–12 mm, significantly longer than sepals;

filaments 1–1.2 mm, anthers 0.4–0.6 mm;

carpels 30–40, apical hairs absent, styles narrowly columnar, distinctly papillate-swollen at very base, 1–1.1 mm.

Achenes

1–1.5 mm, lightly rugose.

1.1–1.5 mm.

Short

hairs not well differentiated from long hairs, absent or sparse throughout.

2n

= 42, 49, 56.

Potentilla wheeleri

Potentilla vahliana

Phenology Flowering May–Sep. Flowering summer.
Habitat Sandy flats, streamsides, lake margins, open conifer woodlands, alpine fellfields Gravelly dry tundra, dry ridges, gravel flats, herb slopes, acidic and calcareous bedrock
Elevation 1800–3500 m (5900–11500 ft) 0–700 m (0–2300 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
NU; QC; Greenland
[BONAP county map]
Discussion

Potentilla wheeleri is found in the southern Sierra Nevada and San Bernardino Mountains of southern California. Compact plants on the summit of Mount San Gorgonio, described by Jepson as var. paupercula, show no consistent difference to justify their taxonomic segregation. Variety viscidula Rydberg has been misapplied to Arizona populations now called P. rhyolitica. Plants identified as P. wheeleri (excluding P. rimicola) from the Sierra San Pedro Mártir (Baja California) stand as a distinct species, P. luteosericea Rydberg (= P. pinetorum Wiggins).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

As circumscribed here, Potentilla vahliana is confined to northeastern arctic Canada (including northernmost Quebec) and western Greenland. Plants previously assigned to P. vahliana from northwestern Greenland, Yukon, most of arctic Canada, and Alaska are now relegated to P. subvahliana.

Plants from western Greenland, whence Potentilla vahliana was described and typified, have petioles with at least some short cottony or crisped hairs in addition to long, smooth hairs, and the styles have distinct papillae at the very base. These features are derived from a combination of the P. uniflora Ledebour and P. nivea groups. Potentilla vahliana is of a higher polyploidy than the species of the P. uniflora group. Potentilla vahliana is treated here as a geographically restricted species, most probably from cross(es) between P. nivea in the narrow sense (probably the Atlantic race, often 2n = 70) and P. subvahliana (2n = 28); it shares several features with the latter, not least of which is leaf dissection and the columnar structure of the caudex.

In mainland Quebec, plants combine features from Potentilla arenosa and P. vahliana; that is, the straight hairs are slightly verrucose, not smooth as in P. nivea, P. subvahliana, and P. vahliana in the narrow sense. They form populations and probably should be described as a new hybrid species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 185. FNA vol. 9, p. 205.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Subviscosae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Niveae
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Synonyms P. wheeleri var. paupercula P. hirsuta
Name authority S. Watson: Proc. Amer. Acad. Arts 11: 148. (1876) Lehmann: Monogr. Potentill., 172. (1820) — not Michaux 1803
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