Potentilla versicolor |
Potentilla villosa |
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Steens Mountain cinquefoil, varying cinquefoil |
hairy cinquefoil, northern cinquefoil, villous cinquefoil |
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Habit | Plants rosetted to ± matted; taproots ± fleshy-thickened. | Plants ± to densely tufted. |
Caudex branches | stout, sometimes short-columnar, not sheathed with marcescent whole leaves. |
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Stems | prostrate to ascending, (0.7–)1.5–2.5(–4) dm, lengths 2–4(–5) times basal leaves. |
ascending, 0.5–2(–2.5) dm, lengths 1.5–3 times basal leaves. |
Basal leaves | pinnate with distal leaflets ± confluent, 4–12 × 1–2.5(–3.5) cm; petiole 1–3 cm, straight hairs absent (on early-season petioles) or sparse to common, spreading-ascending to loosely appressed, 1–2 mm, soft, cottony hairs usually absent, glands sparse; primary lateral leaflets 3–5 per side, on distal (1/3–)1/2–2/3 of leaf axis, overlapping, largest ones cuneate to flabellate, 0.5–1.5(–2) × 0.5–1.5 cm, distal 3/4 to whole margin unevenly incised 2/3 to completely to midvein (blade often medially split as well), ultimate teeth or segments 2–5(–8), ± oblanceolate, 3–11 × 1–3 mm, apical tufts to 1 mm, surfaces green to grayish green, not glaucous, straight hairs sparse to common (sparser adaxially), sometimes absent (except on margins), loosely appressed to ascending, 1–2 mm, soft, cottony hairs absent, glands sparse to common. |
2–12(–15) cm; petiole 1–9(–12) cm, long hairs ± abundant to dense, spreading to ascending, 1–2.5(–3) mm, soft to weak, smooth, crisped hairs absent or sparse, cottony hairs absent, glands sparse to common, sometimes obscured; leaflets usually ± overlapping, central broadly obovate to suborbiculate, (0.5–)1.5–2.5(–3) × (0.5–)1.5–2.6(–3.2) cm, sessile to subsessile, base cuneate to rounded, margins revolute, distal 1/2–2/3(–3/4) incised 1/4–1/2 to midvein, teeth 3–6(–7) per side, ± approximate to ± distant, surfaces ± dissimilar, abaxial yellowish or grayish white, long hairs 1–2 mm, cottony-crisped hairs ± dense, adaxial grayish green, long hairs abundant to dense, crisped hairs absent, sparse, or obscured. |
Cauline leaves | 1–2(–3). |
(0–)1–2. |
Inflorescences | (1–)3–10-flowered, usually openly cymose. |
(1–)2–7(–10)-flowered. |
Pedicels | 1–3(–5) cm, straight to slightly recurved in fruit. |
0.5–3(–4) cm in flower, to 4 cm in fruit. |
Flowers | epicalyx bractlets lanceolate to elliptic, rarely ovate, 2–5 × 1–2 mm, sometimes apically toothed; hypanthium 3–5 mm diam.; sepals 4–7 mm, apex acute; petals 4–7 × 3–5.5 mm; filaments (1–)1.5–2.5 mm, anthers 0.5–0.8 mm; carpels 10–25, styles 2 mm. |
epicalyx bractlets ovate to oval-elliptic, 3–8 × 2–5 mm, 2/3 to as wide as sepals, margins strongly revolute, red glands absent; hypanthium 5–7 mm diam.; sepals 4–8 mm, apex ± acute; petals (5–)7–15 × 7–16 mm, significantly longer than sepals; filaments 1.8–2.1 mm, anthers 0.7–0.8 mm; carpels 150–250, apical hairs usually absent, rarely present (cottony), styles narrowly conic to tapered, ± papillate-swollen on proximal 1/5–1/2, 0.9–1.1 mm. |
Achenes | 1.5–1.8 mm, smooth to faintly rugose, not carunculate. |
0.9–1.3 mm. |
2n | = 14. |
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Potentilla versicolor |
Potentilla villosa |
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Phenology | Flowering summer. | Flowering spring–summer. |
Habitat | Rocky, alpine meadows, seasonally moist slopes, near streams or snowmelt | Sea cliffs, gravel beaches, inland on alpine outcrops (primarily southern populations), pumice barrens, scree and rock ledges |
Elevation | 2100–3200 m (6900–10500 ft) | 0–100(–1800) m (0–300(–5900) ft) |
Distribution |
NV; OR
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AK; OR; WA; BC; e Asia (Russian Far East)
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Discussion | Potentilla versicolor is most common on Steens Mountain in Harney County, Oregon. Collections are known from other mountain ranges in eastern Oregon (some possibly representing distinct taxa) and from near Island Lake in the Ruby Mountains of northeastern Nevada. The species often grows and, apparently, intergrades with P. breweri; it is usually distinct in its lack of cottony hairs. Petals, filaments, and styles tend to be somewhat shorter in P. versicolor than in P. breweri. Collections of P. versicolor have most often been identified as P. breweri, P. millefolia, or P. ovina. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Potentilla villosa is the only documented diploid species of sect. Niveae in North America and, almost certainly, it is fully sexual. It is a characteristic species of coarse-grained beaches and coastal cliffs from southwestern British Columbia to western Alaska and the Russian Far East, and also occurs on scattered mountains in the Olympic Peninsula and Cascade Range of Washington and Oregon (providing the higher elevational extreme). These southern populations, which have been called var. parviflora, tend to be smaller, more delicate, less hairy plants in general, with fewer and smaller flowers. At least some plants in some southern populations also have cottony hairs on the carpels. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 171. | FNA vol. 9, p. 201. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Niveae |
Sibling taxa | ||
Synonyms | P. villosa var. parviflora | |
Name authority | Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 344. (1908) | Pallas ex Pursh: Fl. Amer. Sept. 1: 353. (1813) |
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