Potentilla versicolor |
Potentilla gracilis |
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Steens Mountain cinquefoil, varying cinquefoil |
graceful cinquefoil, northwest cinquefoil, potentille grêle, slender cinquefoil |
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Habit | Plants rosetted to ± matted; taproots ± fleshy-thickened. | |||||||||||||||||
Glands | usually absent or inconspicuous, uncolored. |
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Stems | prostrate to ascending, (0.7–)1.5–2.5(–4) dm, lengths 2–4(–5) times basal leaves. |
ascending to erect, (0.5–)2–7(–12) dm. |
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Basal leaves | pinnate with distal leaflets ± confluent, 4–12 × 1–2.5(–3.5) cm; petiole 1–3 cm, straight hairs absent (on early-season petioles) or sparse to common, spreading-ascending to loosely appressed, 1–2 mm, soft, cottony hairs usually absent, glands sparse; primary lateral leaflets 3–5 per side, on distal (1/3–)1/2–2/3 of leaf axis, overlapping, largest ones cuneate to flabellate, 0.5–1.5(–2) × 0.5–1.5 cm, distal 3/4 to whole margin unevenly incised 2/3 to completely to midvein (blade often medially split as well), ultimate teeth or segments 2–5(–8), ± oblanceolate, 3–11 × 1–3 mm, apical tufts to 1 mm, surfaces green to grayish green, not glaucous, straight hairs sparse to common (sparser adaxially), sometimes absent (except on margins), loosely appressed to ascending, 1–2 mm, soft, cottony hairs absent, glands sparse to common. |
usually not in ranks, palmate, rarely subpalmate, 4–40(–55) cm; petiole (2–)3–25(–45) cm, long hairs sparse to dense, appressed to spreading, 0.5–2(–3) mm, weak to stiff, short hairs absent, crisped hairs usually absent, cottony hairs absent or sparse to abundant (var. owyheensis), glands absent or sparse; leaflets 5–9, on tip, rarely less than distal 1/10, of leaf axis, separate to overlapping, largest ones oblanceolate to elliptic or obovate, (1.5–)2–9(–11) × (0.7–)1.5–4 cm, margins flat to revolute, distal (2/3–)3/4 to nearly whole length evenly to unevenly incised 1/4–3/4+ to midvein, undivided medial blade 2–20 mm wide, teeth (4–)6–10(–11) per side (sometimes secondarily toothed), linear to broadly lanceolate, 2–20(–35) mm, surfaces ± similar to strongly dissimilar, abaxial green or pale green to white, usually not glaucous, long hairs sparse to abundant (sometimes mostly on veins), short-crisped hairs absent or sparse to abundant, cottony hairs absent or sparse to dense, glands usually absent or obscured to sparse, sometimes common, adaxial dark green to grayish, long hairs sparse to ± abundant, rarely absent or dense, short-crisped hairs absent or sparse, cottony hairs absent or sparse to abundant (var. owyheensis), glands usually absent or sparse to sometimes common. |
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Cauline leaves | 1–2(–3). |
1–4(–5). |
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Inflorescences | (1–)3–10-flowered, usually openly cymose. |
(4–)10–50(–60)-flowered. |
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Pedicels | 1–3(–5) cm, straight to slightly recurved in fruit. |
0.3–3 cm. |
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Flowers | epicalyx bractlets lanceolate to elliptic, rarely ovate, 2–5 × 1–2 mm, sometimes apically toothed; hypanthium 3–5 mm diam.; sepals 4–7 mm, apex acute; petals 4–7 × 3–5.5 mm; filaments (1–)1.5–2.5 mm, anthers 0.5–0.8 mm; carpels 10–25, styles 2 mm. |
epicalyx bractlets linear to narrowly lanceolate or elliptic, 2–6(–8) × 0.5–1.5(–2.5) mm, hairs sparse to abundant, rarely dense, ± appressed to ascending, rarely spreading, glands usually absent or inconspicuous; hypanthium 3.5–6 mm diam.; sepals 4–8(–10) mm, apex acute to long acuminate; petals (3–)4–10(–11) × (3–)4–10(–12) mm; filaments (1–)1.5–2.5(–3.5) mm, anthers 0.6–1.2(–1.6) mm; carpels (15–)20–50, styles ± tapered, papillate-swollen proximally, (1–)1.5–2.5(–3) mm. |
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Achenes | 1.5–1.8 mm, smooth to faintly rugose, not carunculate. |
1–1.8 mm. |
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Potentilla versicolor |
Potentilla gracilis |
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Phenology | Flowering summer. | |||||||||||||||||
Habitat | Rocky, alpine meadows, seasonally moist slopes, near streams or snowmelt | |||||||||||||||||
Elevation | 2100–3200 m (6900–10500 ft) | |||||||||||||||||
Distribution |
NV; OR
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AK; AZ; CA; CO; ID; MI; MN; MT; ND; NV; OR; SD; UT; WA; WI; WY; AB; BC; MB; NT; ON; PE; QC; SK; YT
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Discussion | Potentilla versicolor is most common on Steens Mountain in Harney County, Oregon. Collections are known from other mountain ranges in eastern Oregon (some possibly representing distinct taxa) and from near Island Lake in the Ruby Mountains of northeastern Nevada. The species often grows and, apparently, intergrades with P. breweri; it is usually distinct in its lack of cottony hairs. Petals, filaments, and styles tend to be somewhat shorter in P. versicolor than in P. breweri. Collections of P. versicolor have most often been identified as P. breweri, P. millefolia, or P. ovina. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 5 (5 in the flora). Because variation within Potentilla gracilis and other members of sect. Graciles is still far from resolved, taxonomic recognition is given to only the most distinctive extremes and that mostly at the varietal level. The goal is to call attention to areas that need study rather than to provide a full resolution, and even this leaves out some large, widespread populations in both keys and descriptions. The taxonomic placement of some available names, such as P. glomerata A. Nelson, P. longipedunculata Rydberg, and P. macropetala Rydberg, likewise remains unresolved. The vouchers for the report by M. L. Fernald (1950) of Potentilla gracilis and P. pulcherrima from New Hampshire are presumably specimens collected in Coos County by A. S. Pease (NEBC). The three collections differ from one another, corresponding most closely to var. fastigiata, var. flabelliformis, and P. pulcherrima. The presence of such diversity in the same meadow, all as disjunct occurrences, suggests an artificial, non-persisting introduction. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 171. | FNA vol. 9, p. 153. | ||||||||||||||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Graciles | ||||||||||||||||
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Name authority | Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 344. (1908) | Douglas ex Hooker: Bot. Mag. 57: plate 2984. (1830) | ||||||||||||||||
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