Potentilla versicolor |
Potentilla brunnescens |
|
---|---|---|
Steens Mountain cinquefoil, varying cinquefoil |
brownish cinquefoil, slender brown cinquefoil, slender cinquefoil |
|
Habit | Plants rosetted to ± matted; taproots ± fleshy-thickened. | |
Glands | conspicuous, usually golden. |
|
Stems | prostrate to ascending, (0.7–)1.5–2.5(–4) dm, lengths 2–4(–5) times basal leaves. |
ascending to nearly erect, (1.5–)3–7 dm. |
Basal leaves | pinnate with distal leaflets ± confluent, 4–12 × 1–2.5(–3.5) cm; petiole 1–3 cm, straight hairs absent (on early-season petioles) or sparse to common, spreading-ascending to loosely appressed, 1–2 mm, soft, cottony hairs usually absent, glands sparse; primary lateral leaflets 3–5 per side, on distal (1/3–)1/2–2/3 of leaf axis, overlapping, largest ones cuneate to flabellate, 0.5–1.5(–2) × 0.5–1.5 cm, distal 3/4 to whole margin unevenly incised 2/3 to completely to midvein (blade often medially split as well), ultimate teeth or segments 2–5(–8), ± oblanceolate, 3–11 × 1–3 mm, apical tufts to 1 mm, surfaces green to grayish green, not glaucous, straight hairs sparse to common (sparser adaxially), sometimes absent (except on margins), loosely appressed to ascending, 1–2 mm, soft, cottony hairs absent, glands sparse to common. |
sometimes 2-ranked, palmate, (5–)10–30 cm; petiole (1.5–)5–25(–50) cm, long hairs sparse to abundant, appressed to spreading, 1–2 mm, ± stiff, short, crisped, and cottony hairs absent, glands ± abundant; leaflets (5–)7–9, at tip of leaf axis, separate to slightly overlapping, largest ones oblanceolate-elliptic to narrowly obovate, (2–)5–10 × 1–4 cm, margins flat to ± revolute, distal 3/4 to nearly whole length ± evenly incised (1/4–)1/2–3/4+ to midvein, undivided medial blade 2–15(–20) mm wide, teeth (5–)7–10 per side, linear to broadly lanceolate, 3–15 mm, surfaces ± similar, abaxial paler, green, not glaucous, long hairs sparse to common, short-crisped hairs absent or sparse, cottony hairs absent, glands ± abundant. |
Cauline leaves | 1–2(–3). |
1–3(–4). |
Inflorescences | (1–)3–10-flowered, usually openly cymose. |
12–30(–70)-flowered. |
Pedicels | 1–3(–5) cm, straight to slightly recurved in fruit. |
0.2–1(–4) cm. |
Flowers | epicalyx bractlets lanceolate to elliptic, rarely ovate, 2–5 × 1–2 mm, sometimes apically toothed; hypanthium 3–5 mm diam.; sepals 4–7 mm, apex acute; petals 4–7 × 3–5.5 mm; filaments (1–)1.5–2.5 mm, anthers 0.5–0.8 mm; carpels 10–25, styles 2 mm. |
epicalyx bractlets lanceolate to elliptic, 3–6 × 1–2 mm, hairs sparse to common, loosely appressed to spreading, glands common to abundant; hypanthium 4–6(–8) mm diam.; sepals 4–8(–9) mm, apex acute to acuminate; petals 6–9 × 6–10 mm; filaments 1–2 mm, anthers 0.6–1 mm; carpels 30–60, styles filiform-tapered, papillate-swollen proximally, 1.5–2 mm. |
Achenes | 1.5–1.8 mm, smooth to faintly rugose, not carunculate. |
1.5 mm. |
Potentilla versicolor |
Potentilla brunnescens |
|
Phenology | Flowering summer. | Flowering summer. |
Habitat | Rocky, alpine meadows, seasonally moist slopes, near streams or snowmelt | Vernally moist rocky flats, sagebrush benchlands and slopes, usually basaltic |
Elevation | 2100–3200 m (6900–10500 ft) | 400–3100 m (1300–10200 ft) |
Distribution |
NV; OR
|
AK; CO; ID; MT; OR; UT; WA; WY |
Discussion | Potentilla versicolor is most common on Steens Mountain in Harney County, Oregon. Collections are known from other mountain ranges in eastern Oregon (some possibly representing distinct taxa) and from near Island Lake in the Ruby Mountains of northeastern Nevada. The species often grows and, apparently, intergrades with P. breweri; it is usually distinct in its lack of cottony hairs. Petals, filaments, and styles tend to be somewhat shorter in P. versicolor than in P. breweri. Collections of P. versicolor have most often been identified as P. breweri, P. millefolia, or P. ovina. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The conspicuous golden glands, dark green color, and relatively broad leaflets make Potentilla brunnescens one of the better-defined species of sect. Graciles, though it has often been treated as a variety of P. gracilis. The glands, which are often on septate stalks, are particularly conspicuous on the epicalyx bractlets, hypanthium, and sepals, and at the junction of petiole and leaflets. The species occurs from the Blue Mountains of Oregon and Washington to western Montana, northern Utah, and north-central Colorado; collections from Alaska are probably not native. As presented here, Potentilla brunnescens encompasses two morphological extremes that might merit varietal status. Leaflets of the more common and widespread extreme are incised about halfway to the midvein; in contrast, some populations in western Wyoming and adjacent Idaho have leaflets incised 3/4 or more to the midvein. The latter have often been assigned to P. gracilis var. elmeri while reserving P. gracilis var. brunnescens for the former; this is untenable because the type of P. brunnescens is the latter form. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 171. | FNA vol. 9, p. 157. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Graciles |
Sibling taxa | ||
Synonyms | P. gracilis var. brunnescens | |
Name authority | Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 344. (1908) | Rydberg: Bull. Torrey Bot. Club 28: 173. (1901) |
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