Potentilla versicolor
Potentilla biennis
Steens Mountain cinquefoil, varying cinquefoil
biennial cinquefoil, biennial or Greene's cinquefoil, Greene's cinquefoil
prostrate to ascending, (0.7–)1.5–2.5(–4) dm, lengths 2–4(–5) times basal leaves.
ascending to erect, (1–)2–6(–8.5) dm, hairs at base not stiff, not tubercle-based, glands sparse to abundant, conspicuous (to 1 mm, septate).
ternate, (2–)4–12(–25) cm;
petiole (1–)2–8(–20) cm, long hairs sparse to common, spreading to ascending, 0.5–1.5(–3) mm, weak, crisped hairs usually absent, glands sparse to abundant, conspicuous (to 1 mm, septate);
leaflets 3, at tip of leaf axis, usually overlapping, largest ones mostly obovate or oval to nearly round, (0.5–)1–3(–4) × 0.3–3(–3.5) cm, distal 3/4 to ± whole margin evenly to unevenly incised 1/4–1/3(–1/2) to midvein, teeth 3–8(–13) per side, surfaces sparsely to moderately hairy, rarely glabrate, glands sparse to abundant at least abaxially.
pinnate with distal leaflets ± confluent, 4–12 × 1–2.5(–3.5) cm;
petiole 1–3 cm, straight hairs absent (on early-season petioles) or sparse to common, spreading-ascending to loosely appressed, 1–2 mm, soft, cottony hairs usually absent, glands sparse;
primary lateral leaflets 3–5 per side, on distal (1/3–)1/2–2/3 of leaf axis, overlapping, largest ones cuneate to flabellate, 0.5–1.5(–2) × 0.5–1.5 cm, distal 3/4 to whole margin unevenly incised 2/3 to completely to midvein (blade often medially split as well), ultimate teeth or segments 2–5(–8), ± oblanceolate, 3–11 × 1–3 mm, apical tufts to 1 mm, surfaces green to grayish green, not glaucous, straight hairs sparse to common (sparser adaxially), sometimes absent (except on margins), loosely appressed to ascending, 1–2 mm, soft, cottony hairs absent, glands sparse to common.
1–2(–3).
(1–)3–10-flowered, usually openly cymose.
(10–)20–70+-flowered.
1–3(–5) cm, straight to slightly recurved in fruit.
0.2–1(–2) cm.
epicalyx bractlets lanceolate to elliptic, rarely ovate, 2–5 × 1–2 mm, sometimes apically toothed;
hypanthium 3–5 mm diam.;
sepals 4–7 mm, apex acute;
petals 4–7 × 3–5.5 mm;
filaments (1–)1.5–2.5 mm, anthers 0.5–0.8 mm;
carpels 10–25, styles 2 mm.
epicalyx bractlets narrowly oblanceolate to narrowly obovate or elliptic, (1.5–)2–3.5(–4.5) × 0.5–1.5(–2) mm;
hypanthium (2–)3–4(–5.5) mm diam.;
sepals (2–)3–5 mm, apex broadly acute to obtuse;
petals pale yellow to yellow, broadly oblanceolate to obovate, (1–)2–2.7 × 1.5 mm;
stamens 10 or 15, filaments 0.5–1.2 mm, anthers 0.2–0.3 mm;
carpels 40–100, styles 0.5–0.7 mm.
1.5–1.8 mm, smooth to faintly rugose, not carunculate.
whitish or pale yellowish, 0.5–0.9 mm, smooth, without a corky protuberance.
Potentilla versicolor
Potentilla biennis
Potentilla versicolor is most common on Steens Mountain in Harney County, Oregon. Collections are known from other mountain ranges in eastern Oregon (some possibly representing distinct taxa) and from near Island Lake in the Ruby Mountains of northeastern Nevada. The species often grows and, apparently, intergrades with P. breweri; it is usually distinct in its lack of cottony hairs. Petals, filaments, and styles tend to be somewhat shorter in P. versicolor than in P. breweri. Collections of P. versicolor have most often been identified as P. breweri, P. millefolia, or P. ovina.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Potentilla biennis may be recognized by its septate hairs with a round, terminal gland, which dominate the vestiture especially at the base of the plant. Stems are more commonly erect and leaflets rounder than those of P. rivalis. The species is most common along the eastern side of the Sierra Nevada of California, across the Intermountain Region to the Rocky Mountains. All specimens seen by the authors from Baja California, Mexico, which were previously referred to P. biennis are now assigned to P. rivalis.
J. Soják (1996) lectotypified Potentilla millegrana Engelmann ex Lehmann upon a collection of P. rivalis, thus preventing P. biennis from being a later taxonomic synonym.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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