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Potentilla versicolor

Steens Mountain cinquefoil, varying cinquefoil

Habit Plants rosetted to ± matted; taproots ± fleshy-thickened.
Stems

prostrate to ascending, (0.7–)1.5–2.5(–4) dm, lengths 2–4(–5) times basal leaves.

erect, 1–3 dm.

Basal leaves

pinnate with distal leaflets ± confluent, 4–12 × 1–2.5(–3.5) cm;

petiole 1–3 cm, straight hairs absent (on early-season petioles) or sparse to common, spreading-ascending to loosely appressed, 1–2 mm, soft, cottony hairs usually absent, glands sparse;

primary lateral leaflets 3–5 per side, on distal (1/3–)1/2–2/3 of leaf axis, overlapping, largest ones cuneate to flabellate, 0.5–1.5(–2) × 0.5–1.5 cm, distal 3/4 to whole margin unevenly incised 2/3 to completely to midvein (blade often medially split as well), ultimate teeth or segments 2–5(–8), ± oblanceolate, 3–11 × 1–3 mm, apical tufts to 1 mm, surfaces green to grayish green, not glaucous, straight hairs sparse to common (sparser adaxially), sometimes absent (except on margins), loosely appressed to ascending, 1–2 mm, soft, cottony hairs absent, glands sparse to common.

pinnate, (4–)5–10(–14) cm;

petiole 3–7(–10) cm, long hairs abundant, subappresssed to appressed, 0.5–2 mm, ± weak, short hairs absent, crisped or cottony hairs absent or sparse, glands absent or sparse;

leaflets 2–3 per side, on distal 1/3–1/2 of leaf axis, separate, terminal ones elliptic to obovate, (1.8–)2–2.5(–3) × 0.8–1.3 cm, margins revolute, incised nearly to midvein, undivided medial blade 1–2 mm wide, teeth (4–)5–7 per side, linear, surfaces ± dissimilar, abaxial grayish to white, long hairs ± abundant especially on veins, 1–1.5 mm, weak, short hairs absent, crisped hairs abundant to dense, glands absent or obscured, adaxial greenish gray, long hairs common, appressed, 0.5–1 mm, short hairs absent, crisped hairs absent or sparse, glands absent or obscured.

Cauline leaves

1–2(–3).

1–2.

Inflorescences

(1–)3–10-flowered, usually openly cymose.

2–5-flowered, very open.

Pedicels

1–3(–5) cm, straight to slightly recurved in fruit.

2–4 cm.

Flowers

epicalyx bractlets lanceolate to elliptic, rarely ovate, 2–5 × 1–2 mm, sometimes apically toothed;

hypanthium 3–5 mm diam.;

sepals 4–7 mm, apex acute;

petals 4–7 × 3–5.5 mm;

filaments (1–)1.5–2.5 mm, anthers 0.5–0.8 mm;

carpels 10–25, styles 2 mm.

epicalyx bractlets narrowly elliptic to linear, 3–4 × 0.5–0.8(–1) mm, lengths 2/3–3/4 times sepals, margins flat;

hypanthium 2.5–3 mm diam.;

sepals 3.5–5 mm, apex subacute, abaxial surfaces: venation indistinct, glands sparse to common, often obscured;

petals yellow, 7.5–10(–10.5) × 7.5–9(–10) mm, much exceeding sepals;

filaments 1–2 mm, anthers 0.5–0.7 mm;

carpels 50–60, styles papillate-swollen at very base, if at all, 0.8–0.9 mm.

Achenes

1.5–1.8 mm, smooth to faintly rugose, not carunculate.

1.4–1.6 mm, smooth to faintly rugose.

2n

= 28 (14, 42, 56) (Russian Far East).

Potentilla versicolor

Potentilla anachoretica

Phenology Flowering summer. Flowering summer.
Habitat Rocky, alpine meadows, seasonally moist slopes, near streams or snowmelt Dry ridges, rocky outcrops, south-facing slopes, mainly on basic substrates
Elevation 2100–3200 m (6900–10500 ft) 500 m (1600 ft)
Distribution
from FNA
NV; OR
[WildflowerSearch map]
from FNA
AK; e Asia (Russian Far East)
Discussion

Potentilla versicolor is most common on Steens Mountain in Harney County, Oregon. Collections are known from other mountain ranges in eastern Oregon (some possibly representing distinct taxa) and from near Island Lake in the Ruby Mountains of northeastern Nevada. The species often grows and, apparently, intergrades with P. breweri; it is usually distinct in its lack of cottony hairs. Petals, filaments, and styles tend to be somewhat shorter in P. versicolor than in P. breweri. Collections of P. versicolor have most often been identified as P. breweri, P. millefolia, or P. ovina.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Potentilla anachoretica is isolated in sect. Pensylvanicae. B. A. Jurtzev (1993, pers. comm.) claimed indirect support for the presence of P. anachoretica in Alaska and Yukon by the presence of three hybrid species presumably involving P. anachoretica in their parentage: P. borealis Soják, P. murrayi Jurtzev, and P. petrovskyi Soják (sect. Rubricaules; discussed by B. Ertter et al. 2013). Potentilla anachoretica is confirmed from at least two sites in Brooks Range, northern Alaska.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 171. FNA vol. 9, p. 218.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Pensylvanicae
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Name authority Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 344. (1908) Soják: Preslia 46: 70, fig. [p. 71]. (1974)
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