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Mogollon cinquefoil, Navajo cinquefoil

Habit Plants ± tufted.
Caudex branches

slender to stout, not columnar, not sheathed with marcescent whole leaves.

Stems

0.2–1.5(–2) dm.

± ascending, sometimes nearly erect, 0.5–1.5(–3) dm, lengths 2–4 times basal leaves.

Basal leaves

palmate, rarely ternate, (1–)2–7(–18) cm;

petiole (0.5–)1–5(–11) cm, long hairs sparse to abundant, ± spreading (to ascending), 1–3(–4) mm, ± weak, glands ± abundant;

leaflets (3–)5(–7), central flabellate to obovate-cuneate or oblanceolate, (0.5–)1–3(–8) × 0.5–2 cm, not or ± petiolulate, distal 2/3–3/4 of margins evenly to unevenly incised 1/5–3/4 to midvein, sometimes deeply lobed as well, teeth 2–9 per side (some secondarily toothed as well), surfaces green, long hairs sparse to common, 1–2 mm (late-season leaves and adaxial surfaces sometimes nearly glabrate), glands sparse to abundant.

2–4 cm;

petiole 1–2.5 cm, long hairs common to abundant, spreading to ascending, rarely loosely appressed, 1–2 mm, ± stiff, verrucose, short-crisped hairs sparse to abundant, sometimes nearly absent, cottony hairs absent, glands sparse;

leaflets ± overlapping, central elliptic to obovate, 1–1.5 × 0.6–1.1 cm, petiolulate, base broadly cuneate, margins flat or slightly revolute, distal ± 2/3 incised ± 1/2 to midvein, teeth (3–)4 per side, ± approximate, surfaces ± dissimilar, abaxial reddish or greenish gray to white, long hairs 1 mm, cottony-crisped hairs common to dense, adaxial green or reddish to gray-green, long hairs sparse to abundant, short-crisped hairs absent or sparse to common.

Cauline leaves

0–1.

Inflorescences

(1–)3–15-flowered.

2–5-flowered.

Pedicels

0.5–1.5(–2) cm.

1–2 cm in flower, to 3(–5) cm in fruit.

Flowers

epicalyx bractlets lanceolate-elliptic, 1.5–3(–5) × 0.5–1.5 mm;

hypanthium 2.5–4 mm diam.;

sepals (2.5–)3–5(–6) mm, apex ± acute;

petals nearly white abaxially, pale yellow adaxially, narrowly obcordate, 3–6(–8) × 2.5–5 mm;

filaments 1.5–2.5 mm, anthers 0.5–1 mm;

carpels 4–12, styles 2–3 mm.

epicalyx bractlets narrowly elliptic to narrowly ovate, 4–6 × 1–2.5 mm, (1/2–)2/3 to as wide as sepals, margins revolute, sometimes flat, red glands usually common, conspicuous;

hypanthium 3.5–4.5 mm diam.;

sepals 5–7 mm, apex subacute;

petals 6–8 × (4–)5–8 mm, longer than sepals;

filaments 0.7–1.2 mm, anthers 0.4 mm;

carpels 40–60, apical hairs absent, styles columnar, strongly papillate-swollen in proximal 1/5–1/3, 0.9–1 mm.

Achenes

1.5–2 mm, ± rugose.

1.2–1.8 mm.

Short

hairs well differentiated from long hairs, ± abundant to dense throughout.

2n

= 42, 49 (Russian Far East).

Potentilla subviscosa

Potentilla tikhomirovii

Phenology Flowering summer.
Habitat Dry tundra meadows, loamy soil banks, sedge-herb slopes
Elevation 0–1100 m (0–3600 ft)
Distribution
from FNA
AZ; CO; NM
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; NT; NU; Greenland; Europe (Russia); Asia
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Potentilla tikhomirovii is an assumed hybrid species combining characteristics from P. arenosa subsp. arenosa of sect. Niveae (abaxial leaflet surfaces with crisped-cottony hairs, adaxial surfaces with both long and short hairs, petioles often with both long and short hairs, and flower shape and size) and P. hyparctica of sect. Aureae (crisped-cottony hairs often not fully covering abaxial leaf surfaces, reddish color in most plant parts, sepals and epicalyx bractlets relatively large, broad, less acute, and with many and reddish glands, which are often also on petioles and stipules).

Potentilla tikhomirovii has an interrupted range and is not very coherent morphologically. It forms large populations locally, probably by agamic seed propagation, and is a significant part of the Potentilla variation in northern and northeastern Greenland, Ellesmere Island, and parts of the Brooks Range, northern Alaska. Both the distributional and the morphologic patterns indicate that it has evolved multiple times in different regions, such that it is a borderline case for status as an independent species. Its large populations and significant distribution in some regions support recognition, as also does the paucity of obvious back-crosses with the parents.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves often notably seasonally dimorphic; leaflets: early-season evenly to unevenly incised 1/2–3/4 to midvein (sometimes deeply lobed as well), teeth 3–7 per side (sometimes secondarily toothed); late-season evenly incised 1/5–1/4 to midvein, teeth 6–9 per side; petioles: long hairs 1–1.5(–3) mm; n Arizona and New Mexico, barely entering Colorado.
var. subviscosa
1. Leaves not notably seasonally dimorphic; leaflets ± evenly incised 1/4–1/2 to midvein, teeth 2–4(–6 in late-season) per side; petioles: long hairs 2–3(–4) mm; Santa Catalina and Rincon mountains and Sierra Ancha, se Arizona.
var. ramulosa
Source FNA vol. 9, p. 184. FNA vol. 9, p. 201.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Subviscosae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Niveae
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Subordinate taxa
P. subviscosa var. ramulosa, P. subviscosa var. subviscosa
Name authority Greene: Bull. Torrey Bot. Club 8: 97. (1881) Jurtzev: in A. I. Tolmatchew, Fl. Arct. URSS 9(1): 318. (1984)
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