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Mogollon cinquefoil, Navajo cinquefoil

Habit Perennials, rosetted or tufted, not stoloniferous; taproots sometimes ± thickened; vestiture of long and short (rarely short-crisped) hairs, glands sparse to abundant, sometimes red.
Stems

0.2–1.5(–2) dm.

± prostrate to decumbent, sometimes pendent (on rock faces), not flagelliform, not rooting at nodes, lateral to persistent basal rosettes, 0.2–2(–3) dm, lengths 1–3(–5) times basal leaves.

Leaves

basal not in ranks;

cauline 0–2;

primary leaves ternate or palmate, (1–)2–10(–18) cm;

petiole: long hairs usually spreading to ascending, sometimes loosely appressed, weak to stiff, glands usually abundant, sometimes sparse;

leaflets 3–5(–7), at tip of leaf axis, separate to slightly overlapping, oblanceolate or elliptic to broadly obovate or cuneate to flabellate, sometimes nearly orbiculate, margins flat, distal 1/3–3/4 evenly to unevenly (P. subviscosa) incised 1/5–3/4 to midvein, sometimes deeply lobed as well (P. subviscosa), teeth (1–)2–4(–9) per side, surfaces similar, green to gray-green, not glaucous, long hairs weak to stiff, cottony hairs absent.

Basal leaves

palmate, rarely ternate, (1–)2–7(–18) cm;

petiole (0.5–)1–5(–11) cm, long hairs sparse to abundant, ± spreading (to ascending), 1–3(–4) mm, ± weak, glands ± abundant;

leaflets (3–)5(–7), central flabellate to obovate-cuneate or oblanceolate, (0.5–)1–3(–8) × 0.5–2 cm, not or ± petiolulate, distal 2/3–3/4 of margins evenly to unevenly incised 1/5–3/4 to midvein, sometimes deeply lobed as well, teeth 2–9 per side (some secondarily toothed as well), surfaces green, long hairs sparse to common, 1–2 mm (late-season leaves and adaxial surfaces sometimes nearly glabrate), glands sparse to abundant.

Inflorescences

(1–)3–15-flowered.

solitary flowers or 2–10(–20)-flowered, usually ± cymose, open, sometimes racemiform when prostrate.

Pedicels

0.5–1.5(–2) cm.

usually recurved in fruit, 0.5–2(–4) cm, proximal usually not much longer than distal.

Flowers

epicalyx bractlets lanceolate-elliptic, 1.5–3(–5) × 0.5–1.5 mm;

hypanthium 2.5–4 mm diam.;

sepals (2.5–)3–5(–6) mm, apex ± acute;

petals nearly white abaxially, pale yellow adaxially, narrowly obcordate, 3–6(–8) × 2.5–5 mm;

filaments 1.5–2.5 mm, anthers 0.5–1 mm;

carpels 4–12, styles 2–3 mm.

5-merous;

hypanthium 2–6 mm diam.;

petals pale to bright yellow, narrowly or ± obcordate, rarely broadly obovate, (2–)3–7(–8) mm, longer than sepals, apex usually ± retuse;

stamens 15–20;

styles subapical, usually filiform, sometimes tapered-columnar (P. cottamii), papillate-swollen in proximal 1/10(–1/3 in P. cottamii), (1–)1.5–2.5(–3) mm.

Achenes

1.5–2 mm, ± rugose.

smooth to ± rugose.

Short

hairs well differentiated from long hairs, ± abundant to dense throughout.

Potentilla subviscosa

Potentilla sect. Subviscosae

Distribution
from FNA
AZ; CO; NM
[WildflowerSearch map]
[BONAP county map]
sw United States; n Mexico
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species ca. 10 (7 in the flora).

Section Subviscosae consists of species that have radiated in the sky islands of southwestern North America and Mexico, mostly in rocky meadows or outcrops in forested montane areas (B. Ertter 2007c). Some species are chasmophytic, forming clumps on cliffs or steep rock outcrops, similar to species in Ivesia sect. Setosae. Mexican species are poorly known but include at least Potentilla mexiae Standley and possibly P. rydbergiana Rose, as well as undescribed taxa.

Section Subviscosae shares with sect. Concinnae leaves that are often palmate, prostrate stems, and slender pedicels that become recurved in fruit, as well as a similar biogeographic pattern. Section Subviscosae differs in that cottony and strongly verrucose hairs are lacking, leaves are never strongly bicolored, and petiole and stem hairs are spreading to loosely appressed. Vestiture consists of varying proportions of long hairs, short hairs, and glands; petals are often somewhat clawed, unlike the unclawed norm in the genus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves often notably seasonally dimorphic; leaflets: early-season evenly to unevenly incised 1/2–3/4 to midvein (sometimes deeply lobed as well), teeth 3–7 per side (sometimes secondarily toothed); late-season evenly incised 1/5–1/4 to midvein, teeth 6–9 per side; petioles: long hairs 1–1.5(–3) mm; n Arizona and New Mexico, barely entering Colorado.
var. subviscosa
1. Leaves not notably seasonally dimorphic; leaflets ± evenly incised 1/4–1/2 to midvein, teeth 2–4(–6 in late-season) per side; petioles: long hairs 2–3(–4) mm; Santa Catalina and Rincon mountains and Sierra Ancha, se Arizona.
var. ramulosa
1. Basal leaves usually ternate, rarely palmate
→ 2
1. Basal leaves usually palmate, rarely ternate
→ 4
2. Petals (2–)3–4 mm; styles 1–1.5 mm; ne Nevada, nw Utah.
P. cottamii
2. Petals 3–8 mm; styles 2–3 mm; se Arizona
→ 3
3. Petioles: long hairs 1–2(–3) mm; central leaflets with 2–3(–4) teeth per side; Santa Rita, Huachuca, and Chiricahua ranges.
P. rhyolitica
3. Petioles: long hairs 0.5–1(–1.5) mm; central leaflets with 3–5(–6) teeth per side; Pinaleño Mountains, nearby Mogollon Rim, and Santa Catalina Range.
P. albiflora
4. Petals nearly white abaxially, pale yellow adaxially, narrowly obcordate; central leaflets evenly to unevenly incised (sometimes deeply lobed as well), teeth 2–9 per side; short hairs well differentiated from long hairs, ± abundant to dense throughout.
P. subviscosa
4. Petals ± paler abaxially, bright yellow adaxially, ± obcordate; central leaflets evenly incised, teeth 2–4(–5) per side; short hairs not well differentiated from long hairs, absent or sparse throughout
→ 5
5. Plants rooted in ground; leaflets gray-green, long hairs abundant to dense; carpels ca. 20, styles (1.2–)1.5–2 mm; s Sierra Nevada and San Bernardino Mountains, California.
P. wheeleri
5. Plants rooted in rock crevices, often on vertical surfaces; leaflets green, long hairs sparse to abundant; carpels 5–15(–20), styles 1.5–2.5 mm; s California, nw Arizona, and Baja California, Mexico
→ 6
6. Central leaflets 1–3 cm, oblanceolate to broadly obovate-cuneate or nearly orbiculate, distal ± 1/3 of margins incised ± 1/4 to midvein; epicalyx bractlets lanceolate-elliptic, rarely linear; San Jacinto Mountains, California, and Baja California, Mexico.
P. rimicola
6. Central leaflets 0.5–1.5(–2) cm, oblanceolate, distal ± 3/4 of margins incised 1/2–3/4 to midvein; epicalyx bractlets ovate-elliptic; Hualapai Mountains, Arizona.
P. demotica
Source FNA vol. 9, p. 184. FNA vol. 9, p. 182. Author: Barbara Ertter.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Subviscosae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Subordinate taxa
P. subviscosa var. ramulosa, P. subviscosa var. subviscosa
P. albiflora, P. cottamii, P. demotica, P. rhyolitica, P. rimicola, P. subviscosa, P. wheeleri
Synonyms P. unranked Subviscosae
Name authority Greene: Bull. Torrey Bot. Club 8: 97. (1881) (Rydberg) O. Stevens: in N. L. Britton et al., N. Amer. Fl. 22(7): 11. (1959)
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