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Mogollon cinquefoil, Navajo cinquefoil

Hipp's cinquefoil, horse cinquefoil, potentille de Hipp, woolly cinquefoil

Stems

0.2–1.5(–2) dm.

(0.3–)2–4(–5) dm, lengths (1–)2–4 times basal leaves.

Basal leaves

palmate, rarely ternate, (1–)2–7(–18) cm;

petiole (0.5–)1–5(–11) cm, long hairs sparse to abundant, ± spreading (to ascending), 1–3(–4) mm, ± weak, glands ± abundant;

leaflets (3–)5(–7), central flabellate to obovate-cuneate or oblanceolate, (0.5–)1–3(–8) × 0.5–2 cm, not or ± petiolulate, distal 2/3–3/4 of margins evenly to unevenly incised 1/5–3/4 to midvein, sometimes deeply lobed as well, teeth 2–9 per side (some secondarily toothed as well), surfaces green, long hairs sparse to common, 1–2 mm (late-season leaves and adaxial surfaces sometimes nearly glabrate), glands sparse to abundant.

pinnate to subpinnate, (3–)5–15(–25) cm;

petiole 1–10(–15) cm, long hairs abundant to dense, tightly appressed, 1–2.5 mm, stiff, short hairs absent or obscured, crisped-cottony hairs absent or sparse to common, glands absent or obscured;

leaflets not conduplicate, lateral ones evenly (to unevenly in argyrea phase) paired, (2–)3–6(–7) per side on distal (1/6–)1/5–1/2 of leaf axis, distal pairs ± decurrent, often confluent with terminal leaflet, larger leaflets oblanceolate or narrowly obovate to oblong, 1–5(–6) × 0.3–1.5 cm, distal (2/3–)3/4 to whole margin incised 1/2 or less (rarely more) to midvein, teeth (5–)7–12(–18) per side, 1–4 mm, surfaces ± to strongly dissimilar, abaxial white, long hairs abundant (mostly on, but not limited to, veins), 1–2 mm, stiff, short hairs absent or obscured, crisped-cottony hairs abundant to dense, glands absent or obscured, adaxial green to grayish, long hairs sparse to common, rarely absent, short or crisped to, sometimes, cottony hairs absent or sparse to common, glands sparse.

Cauline leaves

1–2(–3).

Inflorescences

(1–)3–15-flowered.

10–30-flowered.

Pedicels

0.5–1.5(–2) cm.

0.3–3(–5) cm.

Flowers

epicalyx bractlets lanceolate-elliptic, 1.5–3(–5) × 0.5–1.5 mm;

hypanthium 2.5–4 mm diam.;

sepals (2.5–)3–5(–6) mm, apex ± acute;

petals nearly white abaxially, pale yellow adaxially, narrowly obcordate, 3–6(–8) × 2.5–5 mm;

filaments 1.5–2.5 mm, anthers 0.5–1 mm;

carpels 4–12, styles 2–3 mm.

epicalyx bractlets narrowly elliptic to lanceolate, rarely linear, 2–5 × 0.5–1.5 mm, (1/2–)2/3 as long as sepals, abaxial vestiture similar to or ± sparser than sepals, usually not glabrescent, straight hairs ± abundant, crisped to sometimes ± cottony hairs absent or sparse to abundant;

hypanthium 3–7 mm diam.;

sepals 4–5.5(–6.5) mm, apex acute to acuminate;

petals 4–8 × 4–7 mm;

filaments 0.5–2.5 mm, anthers 0.6–1.1 mm;

carpels (5–)10–30, styles 1.7–2.5 mm.

Achenes

1.5–2 mm, ± rugose.

1.4–1.8 mm, smooth to faintly rugose.

Short

hairs well differentiated from long hairs, ± abundant to dense throughout.

2n

= 42, 70, 77, 84, 98.

Potentilla subviscosa

Potentilla hippiana

Phenology Flowering summer.
Habitat Dry grasslands and meadows, in aspen and conifer woodlands or alpine tundra, disturbed sites
Elevation 500–3400 m (1600–11200 ft)
Distribution
from FNA
AZ; CO; NM
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; ID; MI; MN; MT; ND; NE; NM; NV; SD; UT; WY; AB; BC; MB; NS; NT; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Potentilla hippiana is most common and distinctive in the Colorado Plateau and southern Rocky Mountains, with outliers in the mountains of Arizona, Nevada, and New Mexico. Occurrences are more scattered in the northern Rocky Mountains and Great Plains, where intergradation with P. effusa is common. Among these intermediates are the argyrea phase and the single report from Idaho (B. C. Johnston 1980). Potentilla hippiana is probably adventive in Nova Scotia, eastern Ontario, Quebec, and Michigan, and possibly the Northwest Territories. The reported occurrence in central Alaska (E. Hultén 1968) is based on a specimen of P. pensylvanica (CAN).

Significant variation exists throughout the range of Potentilla hippiana, including dwarfism, leaf division, leaflet dissection, adaxial leaflet vestiture, and pedicel length. Some of this variation might merit taxonomic recognition upon further analysis, especially if correlated with ploidy level (6x through 12x). At present, the differences used to distinguish var. argyrea are not found to be sufficiently correlated with distribution to merit formal taxonomic recognition.

Complicating the infraspecific variation is a tendency for Potentilla hippiana to intergrade with other species, in particular P. effusa. Beyond sect. Leucophyllae, the frequency of hybrids with P. pulcherrima blurs the distinction between the two species; P. gracilis var. hippianoides S. L. Welsh & N. D. Atwood is probably one of the resultant intermediates.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves often notably seasonally dimorphic; leaflets: early-season evenly to unevenly incised 1/2–3/4 to midvein (sometimes deeply lobed as well), teeth 3–7 per side (sometimes secondarily toothed); late-season evenly incised 1/5–1/4 to midvein, teeth 6–9 per side; petioles: long hairs 1–1.5(–3) mm; n Arizona and New Mexico, barely entering Colorado.
var. subviscosa
1. Leaves not notably seasonally dimorphic; leaflets ± evenly incised 1/4–1/2 to midvein, teeth 2–4(–6 in late-season) per side; petioles: long hairs 2–3(–4) mm; Santa Catalina and Rincon mountains and Sierra Ancha, se Arizona.
var. ramulosa
Source FNA vol. 9, p. 184. FNA vol. 9, p. 162.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Subviscosae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Leucophyllae
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Subordinate taxa
P. subviscosa var. ramulosa, P. subviscosa var. subviscosa
Synonyms P. leucophylla, P. argyrea, P. effusa var. argyrea, P. hippiana var. argyrea, P. hippiana var. diffusa, P. propinqua
Name authority Greene: Bull. Torrey Bot. Club 8: 97. (1881) Lehmann: Nov. Stirp. Pug. 2: 7. (1830) — not Pallas 1773
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