Potentilla subjuga |
Potentilla vulcanicola |
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Colorado cinquefoil |
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Habit | Plants densely tufted. | |
Caudex branches | stout, sometimes ± columnar, not sheathed with marcescent whole leaves. |
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Stems | (0.8–)1–2.5(–3.5) dm. |
ascending to erect, (0.2–)0.4–1.6(–2) dm, lengths 2–3 times basal leaves. |
Basal leaves | usually palmate with additional lateral leaflets, sometimes pinnate, 3–10(–14) cm; petiole 1.5–5 cm, vestiture seasonally dimorphic, long hairs abundant, spreading on first-formed leaves, tightly appressed to ascending on later-formed leaves, 1–2 mm, ± stiff (especially on later-formed leaves), cottony and crisped hairs usually absent, glands absent or sparse; leaflets (3–)5 at tip of leaf axis plus 1(–2) additional pair(s) separated from tip by 3–20 mm, on distal 1/10–1/3(–1/2) of leaf axis, largest leaflets oblanceolate-oblong, (0.5–)1.5–2.5(–3) × 0.3–1 cm, ± whole margin incised 1/2–2/3(–3/4) to midvein, teeth (2–)4–9 per side, usually touching to strongly overlapping, sometimes separate, 2–6 mm, surfaces usually strongly dissimilar (less so on first-formed leaves), abaxial usually white, straight hairs ± abundant (mostly on veins), 1–2 mm, cottony or crisped/cottony hairs ± dense (sparser on first-formed leaves), glands absent or obscured, adaxial green (to grayish), straight hairs sparse to common, 0.5–1.5 mm, cottony and crisped hairs absent, glands sparse. |
1–6(–11) cm; petiole 0.5–4(–9) cm, long hairs ± abundant, ascending to spreading, sometimes loosely appressed, 1–2.5 mm, soft to weak, smooth, crisped(/cottony) hairs absent or sparse, sometimes common, glands absent or sparse; leaflets overlapping, central broadly obovate, 0.8–2(–2.5) × 0.8–1.5(–2.5) cm, sessile or subsessile, base broadly cuneate, margins strongly revolute, distal 1/2–2/3 incised ± 1/2 to midvein, teeth 2–3(–4) per side, distant, surfaces strongly dissimilar, abaxial grayish white, long hairs 1–2 mm, cottony-crisped hairs ± dense, adaxial usually dark green, sometimes grayish green, long hairs sparse to abundant, other hairs usually absent. |
Cauline leaves | 1–3. |
(0–)1–2(–3). |
Inflorescences | 3–20(–30)-flowered. |
1–2(–5)-flowered. |
Pedicels | 0.5–2 cm (proximal to 3 cm). |
1–2 cm in flower, to 5 cm in fruit. |
Flowers | epicalyx bractlets narrowly to broadly lanceolate, 2–5(–6) × 1–1.5 mm; sepals 4–7 mm, apex acute to acuminate; petals 4–8 × 4–8 mm; filaments (0.5–)1–2 mm, anthers 0.3–0.8 mm; carpels 15–30, styles filiform to filiform-tapered, ± papillate-swollen in less than proximal 1/5, 1.5–2 mm. |
epicalyx bractlets ± oblong to ovate, 4–8(–10) × 1.8–3(–5) mm, 2/3 to as wide as sepals, margins revolute, sometimes flat, red glands absent; hypanthium 3.5–4.5 mm diam.; sepals 5–8 mm, apex ± acute; petals 7–9 × 7–10 mm, longer than sepals; filaments 0.8–1 mm, anthers 0.6–0.7 mm; carpels 20–30, apical hairs sparse to abundant (straight), styles conic-columnar, ± papillate-swollen in less than proximal 1/5, or sometimes to 1/5, 0.8–1.1 mm. |
Achenes | 1.2–1.6 mm. |
0.9–1.2 mm. |
2n | = 28 (Russian Far East). |
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Potentilla subjuga |
Potentilla vulcanicola |
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Phenology | Flowering summer. | Flowering late spring to summer. |
Habitat | Alpine tundra and meadows, boulder piles, gravelly slopes, stabilized talus | Dry alpine heaths, ridge crests, rock outcrops, herb slopes, scree and talus, stabilized sand dunes, mainly on calcareous bedrock |
Elevation | 3400–4000 m (11200–13100 ft) | 0–3100 m (0–10200 ft) |
Distribution |
CO; NM; AB
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AK; NT; NU; YT; e Asia (Russian Far East) |
Discussion | Potentilla subjuga is centered in the high mountains of Colorado and barely enters New Mexico in the Sangre de Cristo Mountains. Collections from Alberta also apparently belong to this species, but all known collections from Wyoming have been identified as different taxa, at least one currently undescribed. At its most distinctive, P. subjuga is easily recognized by its unique leaf division, with five palmate leaflets subtended by an additional pair (or two) of lateral leaflets. Southern populations, however, are more likely to have only three apical leaflets. The leaflets tend to be strongly bicolored with overlapping teeth, in contrast to most sympatric pinnate species. Petiole vestiture is also distinctive in being seasonally dimorphic, with long hairs on first-formed leaves spreading to ascending and those on later formed leaves tightly appressed, as well as more conspicuously verrucose. Unresolved infraspecific variation exists, and field observations suggest that P. subjuga readily hybridizes with sympatric species, creating a swarm of intermediate specimens. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Potentilla vulcanicola is close morphologically to P. uniflora in the narrow sense and largely replaces it in southern and eastern Chukotka, Alaska, and northwestern Canada, where it reaches east of Mackenzie River on the coastal mainland and the westernmost islands (Banks and Victoria islands). Potentilla subgorodkovii and P. vulcanicola account for a major part of North American records of P. uniflora. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 166. | FNA vol. 9, p. 203. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Subjugae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Niveae |
Sibling taxa | ||
Synonyms | P. osterhoutiana | |
Name authority | Rydberg: Bull. Torrey Bot. Club 23: 397, plate 274. (1896) | Juzepczuk: Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 17: 222. (1955) |
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