Potentilla subjuga
Potentilla uliginosa
Colorado cinquefoil
Cunningham Marsh cinquefoil
(0.8–)1–2.5(–3.5) dm.
probably prostrate to decumbent, or ± ascending in supporting vegetation, 2.5–5.5 dm, lengths 1–2 times basal leaves.
usually palmate with additional lateral leaflets, sometimes pinnate, 3–10(–14) cm;
petiole 1.5–5 cm, vestiture seasonally dimorphic, long hairs abundant, spreading on first-formed leaves, tightly appressed to ascending on later-formed leaves, 1–2 mm, ± stiff (especially on later-formed leaves), cottony and crisped hairs usually absent, glands absent or sparse;
leaflets (3–)5 at tip of leaf axis plus 1(–2) additional pair(s) separated from tip by 3–20 mm, on distal 1/10–1/3(–1/2) of leaf axis, largest leaflets oblanceolate-oblong, (0.5–)1.5–2.5(–3) × 0.3–1 cm, ± whole margin incised 1/2–2/3(–3/4) to midvein, teeth (2–)4–9 per side, usually touching to strongly overlapping, sometimes separate, 2–6 mm, surfaces usually strongly dissimilar (less so on first-formed leaves), abaxial usually white, straight hairs ± abundant (mostly on veins), 1–2 mm, cottony or crisped/cottony hairs ± dense (sparser on first-formed leaves), glands absent or obscured, adaxial green (to grayish), straight hairs sparse to common, 0.5–1.5 mm, cottony and crisped hairs absent, glands sparse.
pinnate with distal leaflets ± confluent, 15–32 × 2–4 cm;
petiole 5–10(–15) cm, straight hairs absent or sparse to common, appressed, 0.5–1.5 mm, stiff, cottony hairs absent, glands absent or sparse;
primary lateral leaflets 6–10(–12) per side (irregularly paired), on distal ± 1/2 of leaf axis, loosely overlapping to proximally separate, largest ones cuneate to flabellate, 1.2–2.2 × 1–3 cm, ± whole margin unevenly incised 3/4 to nearly to midvein, ultimate segments (3–)5–10(–15), linear to narrowly oblanceolate, 5–17 × 1–2.5 mm, apical tufts less than 0.5 mm, surfaces green, not glaucous, straight hairs sparse (often glabrate adaxially), appressed, 0.5–1(–1.5) mm, stiff, cottony hairs absent, glands absent or sparse.
1–3.
2.
3–20(–30)-flowered.
6–10-flowered, very openly cymose.
0.5–2 cm (proximal to 3 cm).
1–6 cm, ± recurved in fruit.
epicalyx bractlets narrowly to broadly lanceolate, 2–5(–6) × 1–1.5 mm;
sepals 4–7 mm, apex acute to acuminate;
petals 4–8 × 4–8 mm;
filaments (0.5–)1–2 mm, anthers 0.3–0.8 mm;
carpels 15–30, styles filiform to filiform-tapered, ± papillate-swollen in less than proximal 1/5, 1.5–2 mm.
epicalyx bractlets narrowly elliptic to oblong-ovate, 4–6 × 1–2.5 mm;
hypanthium 5–6 mm diam.;
sepals 4–7 mm, apex acute;
petals 6–10 × 5–8 mm;
filaments (1.5–)2–3 mm, anthers 0.7–1.2 mm;
carpels ca. 10, styles 2.5–3.5 mm.
1.2–1.6 mm.
2–2.6 mm, smooth, ± carunculate.
Potentilla subjuga
Potentilla uliginosa
Potentilla subjuga is centered in the high mountains of Colorado and barely enters New Mexico in the Sangre de Cristo Mountains. Collections from Alberta also apparently belong to this species, but all known collections from Wyoming have been identified as different taxa, at least one currently undescribed. At its most distinctive, P. subjuga is easily recognized by its unique leaf division, with five palmate leaflets subtended by an additional pair (or two) of lateral leaflets. Southern populations, however, are more likely to have only three apical leaflets. The leaflets tend to be strongly bicolored with overlapping teeth, in contrast to most sympatric pinnate species. Petiole vestiture is also distinctive in being seasonally dimorphic, with long hairs on first-formed leaves spreading to ascending and those on later formed leaves tightly appressed, as well as more conspicuously verrucose. Unresolved infraspecific variation exists, and field observations suggest that P. subjuga readily hybridizes with sympatric species, creating a swarm of intermediate specimens.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Of conservation concern.
Potentilla uliginosa is known only from a handful of historical collections from Cunningham Marsh, one of three wetlands in southern Sonoma County that harbor a suite of disjunct and regionally rare species (B. C. Johnston and B. Ertter 2010). Although previously included in P. hickmanii, P. uliginosa differs in having woodier caudices, longer stems and leaves, and more deeply incised leaflets occupying less of the leaf axis. Potentilla uliginosa is presumed extinct; however, a morphologically comparable collection of the P. millefolia complex from Josephine County, Oregon (Deer Creek, near Selma, 14 June 1929, Applegate 5735, UC), opens the possibility of additional undiscovered populations in the mountains of northwestern California.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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