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Colorado cinquefoil

scarlet cinquefoil, Thurber's cinquefoil

Stems

(0.8–)1–2.5(–3.5) dm.

(2–)3–7(–10) dm.

Basal leaves

usually palmate with additional lateral leaflets, sometimes pinnate, 3–10(–14) cm;

petiole 1.5–5 cm, vestiture seasonally dimorphic, long hairs abundant, spreading on first-formed leaves, tightly appressed to ascending on later-formed leaves, 1–2 mm, ± stiff (especially on later-formed leaves), cottony and crisped hairs usually absent, glands absent or sparse;

leaflets (3–)5 at tip of leaf axis plus 1(–2) additional pair(s) separated from tip by 3–20 mm, on distal 1/10–1/3(–1/2) of leaf axis, largest leaflets oblanceolate-oblong, (0.5–)1.5–2.5(–3) × 0.3–1 cm, ± whole margin incised 1/2–2/3(–3/4) to midvein, teeth (2–)4–9 per side, usually touching to strongly overlapping, sometimes separate, 2–6 mm, surfaces usually strongly dissimilar (less so on first-formed leaves), abaxial usually white, straight hairs ± abundant (mostly on veins), 1–2 mm, cottony or crisped/cottony hairs ± dense (sparser on first-formed leaves), glands absent or obscured, adaxial green (to grayish), straight hairs sparse to common, 0.5–1.5 mm, cottony and crisped hairs absent, glands sparse.

palmate, (2–)4–15(–30) cm;

petiole (2–)3–10(–25) cm, long hairs sparse to common, spreading, 1–3.5 mm, weak, glands sparse to abundant, sometimes absent;

leaflets 5(–7), at tip of leaf axis, central one (1–)2–6(–8) × (0.5–)1–2.5(–4) cm, petiolules 0(–10) mm, distal 3/4 to whole margin incised 1/5–1/4 to midvein, teeth (4–)7–15(–19) per side (sometimes secondarily toothed), (1–)2–3(–5) mm, teeth apex acute to ± obtuse, surfaces slightly to strongly dissimilar, abaxial pale green to white, straight hairs sparse to abundant, often dense on veins, 0.5–3 mm, cottony hairs absent or sparse to dense, glands ± sparse or absent, sometimes obscured, adaxial green, straight hairs sparse to abundant, 0.2–1 mm, cottony hairs absent, glands ± sparse or absent.

Cauline leaves

1–3.

stipules usually toothed, sometimes entire.

Inflorescences

3–20(–30)-flowered.

(4–)10–35-flowered.

Pedicels

0.5–2 cm (proximal to 3 cm).

0.5–4 cm.

Flowers

epicalyx bractlets narrowly to broadly lanceolate, 2–5(–6) × 1–1.5 mm;

sepals 4–7 mm, apex acute to acuminate;

petals 4–8 × 4–8 mm;

filaments (0.5–)1–2 mm, anthers 0.3–0.8 mm;

carpels 15–30, styles filiform to filiform-tapered, ± papillate-swollen in less than proximal 1/5, 1.5–2 mm.

epicalyx bractlets narrowly to broadly lanceolate-elliptic (rarely toothed or lobed), 4–9(–14) × (1–)1.5–2 mm;

sepals 4–10(–15) mm, apex acute to acuminate;

petals ± dark reddish throughout, (3.5–)6–10 × 6–10 mm;

filaments 1.5–3 mm, anthers 0.8–1 mm;

carpels 30–70, styles 2.5–3.5 mm.

Achenes

1.2–1.6 mm.

1.5 mm, ± rugose.

Potentilla subjuga

Potentilla thurberi

Phenology Flowering summer.
Habitat Alpine tundra and meadows, boulder piles, gravelly slopes, stabilized talus
Elevation 3400–4000 m (11200–13100 ft)
Distribution
from FNA
CO; NM; AB
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Potentilla subjuga is centered in the high mountains of Colorado and barely enters New Mexico in the Sangre de Cristo Mountains. Collections from Alberta also apparently belong to this species, but all known collections from Wyoming have been identified as different taxa, at least one currently undescribed. At its most distinctive, P. subjuga is easily recognized by its unique leaf division, with five palmate leaflets subtended by an additional pair (or two) of lateral leaflets. Southern populations, however, are more likely to have only three apical leaflets. The leaflets tend to be strongly bicolored with overlapping teeth, in contrast to most sympatric pinnate species. Petiole vestiture is also distinctive in being seasonally dimorphic, with long hairs on first-formed leaves spreading to ascending and those on later formed leaves tightly appressed, as well as more conspicuously verrucose. Unresolved infraspecific variation exists, and field observations suggest that P. subjuga readily hybridizes with sympatric species, creating a swarm of intermediate specimens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

The abaxially white-cottony leaflets of var. atrorubens usually make it easily distinguished from var. thurberi, which lacks cottony hairs and is usually less hairy in general. The two varieties sometimes occur together and intermediates between the extremes are common.

The degree of glandularity and the lengths of the central leaflets are greater in the southern populations of both varieties. The glands are less widely distributed on the plants northward and are fewer in northern Arizona and New Mexico. Leaflets are shorter and broader north of Graham County, Arizona, and Otero County, New Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaflets pale green abaxially, cottony hairs absent.
var. thurberi
1. Leaflets ± white to white abaxially, cottony hairs sparse to dense.
var. atrorubens
Source FNA vol. 9, p. 166. FNA vol. 9, p. 149.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Subjugae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Rubrae
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Subordinate taxa
P. thurberi var. atrorubens, P. thurberi var. thurberi
Synonyms P. osterhoutiana
Name authority Rydberg: Bull. Torrey Bot. Club 23: 397, plate 274. (1896) A. Gray: Pl. Nov. Thurb., 318. (1854)
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