Potentilla subjuga
Potentilla subvahliana
Colorado cinquefoil
high arctic cinquefoil
stout, columnar, sheathed with marcescent whole leaves.
(0.8–)1–2.5(–3.5) dm.
erect, 0.2–0.6(–1.1) dm, lengths 2–4 times basal leaves.
usually palmate with additional lateral leaflets, sometimes pinnate, 3–10(–14) cm;
petiole 1.5–5 cm, vestiture seasonally dimorphic, long hairs abundant, spreading on first-formed leaves, tightly appressed to ascending on later-formed leaves, 1–2 mm, ± stiff (especially on later-formed leaves), cottony and crisped hairs usually absent, glands absent or sparse;
leaflets (3–)5 at tip of leaf axis plus 1(–2) additional pair(s) separated from tip by 3–20 mm, on distal 1/10–1/3(–1/2) of leaf axis, largest leaflets oblanceolate-oblong, (0.5–)1.5–2.5(–3) × 0.3–1 cm, ± whole margin incised 1/2–2/3(–3/4) to midvein, teeth (2–)4–9 per side, usually touching to strongly overlapping, sometimes separate, 2–6 mm, surfaces usually strongly dissimilar (less so on first-formed leaves), abaxial usually white, straight hairs ± abundant (mostly on veins), 1–2 mm, cottony or crisped/cottony hairs ± dense (sparser on first-formed leaves), glands absent or obscured, adaxial green (to grayish), straight hairs sparse to common, 0.5–1.5 mm, cottony and crisped hairs absent, glands sparse.
0.5–2.5(–3) cm;
petiole 0.3–1.5(–2) cm, long hairs common to abundant, spreading to ascending, 1(–2) mm, ± weak, rarely stiff, smooth, crisped/short-cottony hairs usually absent, sometimes sparse, glands sparse to common;
leaflets separate to slightly overlapping, central obovate, 0.5–1.2(–1.5) × 0.4–0.9(–1) cm, subsessile to short-petiolulate, base cuneate, margins revolute, distal 1/2 incised 1/2–3/4 to midvein, teeth (1–)2–3(–4) per side, ± distant, surfaces ± dissimilar, abaxial grayish or yellowish white, long hairs 0.5–1 mm, cottony-crisped hairs ± dense (often obscured by long hairs), adaxial dark green to greenish gray, long hairs ± abundant, other hairs absent.
1–3.
(0–)1(–2).
3–20(–30)-flowered.
1(–2)-flowered.
0.5–2 cm (proximal to 3 cm).
1–2(–3) cm in flower, to 5.5 cm in fruit.
epicalyx bractlets narrowly to broadly lanceolate, 2–5(–6) × 1–1.5 mm;
sepals 4–7 mm, apex acute to acuminate;
petals 4–8 × 4–8 mm;
filaments (0.5–)1–2 mm, anthers 0.3–0.8 mm;
carpels 15–30, styles filiform to filiform-tapered, ± papillate-swollen in less than proximal 1/5, 1.5–2 mm.
epicalyx bractlets lanceolate to elliptic-ovate, rarely linear, 3–6(–7) × 0.8–2(–2.3) mm, (1/2–)2/3 to nearly as wide as sepals, margins flat or ± revolute, red glands absent;
hypanthium 2.5–4 mm diam.;
sepals 3–5(–6) mm, apex obtuse to subacute;
petals 4–8(–9) × 4–9 mm, longer than sepals;
filaments 1–1.3 mm, anthers 0.3–0.5 mm;
carpels 25–35, apical hairs absent, styles conic-columnar, ± papillate-swollen on less than proximal 1/5, 0.9–1.1 mm.
1.2–1.6 mm.
1.2–1.6 mm.
= 28 (Russian Far East).
Potentilla subjuga
Potentilla subvahliana
Potentilla subjuga is centered in the high mountains of Colorado and barely enters New Mexico in the Sangre de Cristo Mountains. Collections from Alberta also apparently belong to this species, but all known collections from Wyoming have been identified as different taxa, at least one currently undescribed. At its most distinctive, P. subjuga is easily recognized by its unique leaf division, with five palmate leaflets subtended by an additional pair (or two) of lateral leaflets. Southern populations, however, are more likely to have only three apical leaflets. The leaflets tend to be strongly bicolored with overlapping teeth, in contrast to most sympatric pinnate species. Petiole vestiture is also distinctive in being seasonally dimorphic, with long hairs on first-formed leaves spreading to ascending and those on later formed leaves tightly appressed, as well as more conspicuously verrucose. Unresolved infraspecific variation exists, and field observations suggest that P. subjuga readily hybridizes with sympatric species, creating a swarm of intermediate specimens.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
The major part of the range assigned to Potentilla vahliana by E. Hultén (1968) belongs to P. subvahliana.
Two morphologic types are present within what is accepted here as Potentilla subvahliana. Plants corresponding to the type of P. subvahliana (Wrangel Island) are widespread throughout northeastern Asia (Chukotka) and arctic North America to northwestern Greenland. Some plants in alpine Alaska, Yukon, and east to Amundsen Gulf, Nunavut, are redder, more densely columnar, and less hairy, with smaller leaves having fewer and narrower lobes, more slender one-flowered stems, narrow and entire bracts, narrower sepals, and much narrower epicalyx bractlets. Potentilla subvahliana is tetraploid, perhaps an allopolyploid, and could consist of lineages from different parental combinations.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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