Potentilla subgorodkovii |
Potentilla bimundorum |
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Beringian cinquefoil, yurtsev's cinquefoil |
cutleaf cinquefoil, potentille des deux mondes, staghorn cinquefoil |
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Habit | Plants ± densely tufted. | |
Caudex branches | stout, usually columnar, sometimes sheathed with marcescent whole leaves. |
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Stems | erect, (0.2–)0.3–1.5(–2) dm, lengths 1.5–2.5(–3.5) times basal leaves. |
decumbent to ascending, rarely erect, 1–5 dm. |
Basal leaves | (1–)2–10(–15) cm; petiole (0.5–)1–5(–12) cm, long hairs common to dense, ± ascending to loosely appressed, sometimes spreading, 1–2 mm, ± soft, smooth, crisped/short-cottony hairs usually sparse, sometimes absent or common, glands absent, sparse, or obscured; leaflets separate to slightly overlapping, central obovate or obtriangular, (0.5–)1–2(–3) × (0.4–)0.8–1.5(–2) cm, sessile or subsessile, base cuneate, margins revolute, distal (1/3–)1/2–2/3(–3/4) incised 1/2–2/3(–3/4) to midvein, teeth 2–3(–4) per side, usually ± distant, surfaces somewhat to more often strongly dissimilar, abaxial yellowish or grayish white to white, long hairs 0.8–1.5 mm, cottony-crisped hairs ± dense, adaxial green to grayish green, long hairs sparse to ± abundant, other hairs usually absent. |
subpinnate, 7–22(–25) cm; petiole 3–12(–15) cm, long hairs ± abundant, appressed to ascending, 0.5–3 mm, stiff, short hairs absent, cottony or crisped hairs absent, glands absent or sparse; leaflets 3(–4) per side, on distal 1/5–1/3(–1/2) of leaf axis, overlapping or not, terminal ones oblong to broadly oblong, 1.5–5 × 0.7–2 cm, margins strongly revolute, incised nearly to midvein, undivided medial blade 1–3 mm wide, teeth (3–)4–5 per side, linear, rarely narrowly oblong to narrowly lanceolate, surfaces strongly dissimilar, abaxial white or grayish white, long hairs abundant, ± 1 mm, stiff verrucose on veins, sparse and soft between veins, short hairs absent, crisped and cottony hairs dense, glands absent, adaxial dark green, long hairs sparse to common, tightly appressed, 0.5–1 mm, short hairs absent, cottony and crisped hairs absent, glands absent or sparse. |
Cauline leaves | (0–)1(–2). |
(1–)2–3. |
Inflorescences | usually 1(–2)-flowered, rarely to 5-flowered. |
(5–)10–40+-flowered, ± congested to open. |
Pedicels | (0.5–)2–4 cm in flower, to 5 cm in fruit. |
0.3–1 cm (proximal to 3 cm). |
Flowers | epicalyx bractlets elliptic-lanceolate to ovate, (3–)4–6(–7) × (0.8–)1.2–2(–2.5) mm, (1/2–)2/3 to as wide as sepals, margins revolute, rarely flat, red glands absent; hypanthium 2.5–4 mm diam.; sepals 4–6(–7) mm, apex subacute; petals (5–)6–9 × (5–)7–9 mm, significantly longer than sepals; filaments 1–1.3 mm, anthers 0.4–0.6 mm; carpels 30–40, apical hairs absent, styles narrowly columnar to conic-tapered, papillate-swollen on proximal 1/5(–1/3), 0.9–1.1 mm. |
epicalyx bractlets linear to narrowly oblong-lanceolate, 3–5 × 0.9–1.3 mm, lengths ± 3/4 times sepals, margins flat; hypanthium 2.5–3.5 mm diam.; sepals 4–7(–8) mm, apex acute to obtuse, abaxial surfaces: venation prominent at least proximally, glands ± sparse, obscured to evident; petals pale yellow, 3.5–5 × 3–5 mm, lengths ± equal to sepals; filaments 0.5–1.5 mm, anthers 0.5–0.6 mm; carpels 40–60, styles papillate-swollen at very base, if at all, 0.8–0.9 mm. |
Achenes | 1.2–2 mm. |
1.3–1.5 mm, rugose. |
2n | = 28, 42, 49, 56 (Russian Far East). |
= 28. |
Potentilla subgorodkovii |
Potentilla bimundorum |
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Phenology | Flowering late spring to summer. | Flowering summer. |
Habitat | Dry alpine heaths, exposed ridges and summits, rock outcrops, scree and talus, dry tundra, acidic and calcareous bedrock | Road and railway verges, gravelly ruderal sites, acidic rocky outcrops, stream banks and gravel bars, sandy-gravelly flood plains, gravel ridges, sandy lakeshores, sandy steppe bluffs |
Elevation | 0–4300 m (0–14100 ft) | 0–1000 m (0–3300 ft) |
Distribution |
AK; CO; MT; UT; WY; AB; BC; NT; NU; YT; Asia (Russian Far East, Sakha [Yakutia]) |
AK; AB; MB; NT; NU; ON; QC; SK; YT; e Asia (Russian Far East) |
Discussion | The name Potentilla subgorodkovii was coined for a presumed hybrid species resulting from P. crebridens × P. subvahliana; here it is applied in a collective meaning for plants combining characteristics from multiple species of the P. uniflora/villosa and P. nivea groups. Morphologically, this collective entity is much closer to the P. uniflora/villosa group than to the P. nivea group and is clearly different from primary hybrids and clones, which are often observed. Potentilla subgorodkovii constitutes about half of what previously has been considered P. uniflora in Alaska and Yukon and is the only such entity fully confirmed south of northern British Columbia. These southern populations occur outside the range of possible parental members of the P. uniflora/villosa group and do not support the hybrid hypothesis. They may instead represent a distinct species, not yet described. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Potentilla bimundorum is the correct name for what has been treated as P. multifida Linnaeus in North America. It is the only American representative of the mainly Eurasian P. multifida group, which is sometimes treated as a distinct section. J. Soják (2005) presented a useful survey and key to this group, showing that its species, in spite of being merged within P. multifida in the broad sense by many authors, are each distinct and with their own range. Potentilla multifida in the narrow sense is restricted to eastern Europe and western Siberia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 204. | FNA vol. 9, p. 218. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Niveae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Pensylvanicae |
Sibling taxa | ||
Synonyms | P. multifida subsp. bimundorum | |
Name authority | Jurtzev: Bot. Zhurn. (Moscow & Leningrad) 78: 83. (1993) | Soják: Č as. Nár. Muz. Praze Rada P ř ír. 141: 195. (1974) |
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