Potentilla sect. Subjugae
Rosaceae subfam. rosoideae
ascending to nearly erect, not flagelliform, not rooting at nodes, lateral to persistent basal rosettes, (0.2–)0.4–2.5(–3.5) dm, lengths 2–3 times basal leaves.
basal not 2-ranked;
cauline 0–3;
primary leaves palmate with additional lateral pairs to pinnate (with distal leaflets distinct), (1–)1.5–10(–14) cm;
petiole: long hairs spreading to tightly appressed, weak to stiff, glands absent or sparse;
leaflets 5–7(–9), on distal 1/10–1/3(–1/2) of leaf axis, strongly overlapping or not, oblanceolate-oblong to obovate, margins scarcely to ± revolute, ± whole length evenly incised 1/2–3/4 to midvein, teeth 2–9 per side, surfaces ± to strongly dissimilar, abaxial white to grayish green, cottony hairs sparse to dense, adaxial green to grayish, not glaucous, long hairs ± stiff.
alternate, rarely opposite, pinnately compound, sometimes simple or palmately compound;
stipules present, rarely absent.
(1–)2–20(–30)-flowered, usually cymose, open.
straight in fruit, 0.5–2(–3) cm, proximal ± longer than distal.
5-merous;
hypanthium 3–5 mm diam.;
petals yellow, ± obcordate, 4–8 mm, equal to or longer than sepals, apex retuse;
stamens ca. 20;
styles subapical, filiform to filiform-tapered or filiform-columnar, papillate-swollen in proximal less than 1/5 if at all, 0.8–2 mm.
torus usually enlarged, sometimes small or absent;
carpels 1–260(–450), distinct, free, styles distinct, rarely connate (Roseae);
ovules 1(or 2), collateral (Rubeae) or superposed (Fallugia, Filipendula).
achenes or aggregated achenes sometimes with fleshy, urn-shaped hypanthium or enlarged torus, sometimes aggregated drupelets;
styles persistent or deciduous, not elongate (elongate but not plumose in Geum).
smooth.
= 7(8).
Potentilla sect. Subjugae
Rosaceae subfam. rosoideae
Species 2 (2 in the flora).
Section Subjugae is used primarily to accommodate the unique leaf dissection and vestiture of Potentilla subjuga, which commonly has five palmately arranged leaflets and one, sometimes two, additional pairs of lateral leaflets. Sometimes leaves are simply pinnate with only three terminal leaflets, and sufficient intergradation occurs with P. saximontana that it is also included in the section. This circumscription coincides with the initial concept of Rydberg and Nelson and contrasts with that of B. C. Johnston (1980, 1985).
Section Subjugae is restricted to subalpine and alpine habitats in the Rocky Mountains and adjacent ranges of western North America centered in Colorado. Variation within the section is complex and poorly understood, such that the two species treated here represent only the best expressed extremes with existing names. Other named components of uncertain placement include Potentilla lupina Rydberg, P. minutifolia Rydberg, P. rubricaulis var. nana Clements & E. G. Clements, P. rubripes Rydberg [= P. concinna var. rubripes (Rydberg) C. L. Hitchcock], and P. tenerrima Rydberg.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Variation in the number of genera in subfam. Rosoideae is due to differences in generic delimitation between D. Potter et al. (2007) and the authors of some Potentilleae genera. Cyanogenic glycosides and sorbitol are absent in the subfamily.
Tribes 6, genera 28–35, species ca. 1600 (6 tribes, 26 genera, 302 species, including 1 hybrid, in the flora)
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Leaflets at tip of leaf axis (3–)5; styles 1.5–2 mm; stems (0.8–)1–2.5(–3.5) dm; inflorescences 3–20(–30)-flowered. | P. subjuga |
1. Leaflets at tip of leaf axis 3; styles 0.8–1.5 mm; stems (0.2–)0.4–1(–1.5) dm; inflorescences (1–)2–5(–10)-flowered. | P. saximontana |
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