Potentilla sect. Aureae |
Potentilla grayi |
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Gray's cinquefoil |
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Habit | Perennials, tufted to cushion-forming or matted, rarely rosetted, not stoloniferous (except sometimes P. verna); taproots not fleshy-thickened; vestiture primarily of straight hairs, not differentiated into long and short, glands absent or sparse to abundant, sometimes red (P. hyparctica). | Plants rosetted or ± tufted; caudex branches absent or short to elongate, stout to slender. | ||||||||||||||||||||||||||||||||||||||||||||||||
Stems | usually ascending to erect, sometimes prostrate to decumbent or spreading, not flagelliform, not rooting at nodes (except P. verna), lateral to persistent basal rosettes, 0.1–2(–3) dm, lengths 1–3(–4) times basal leaves. |
ascending to erect, (0.4–)1–2 dm, lengths 2–4 times basal leaves. |
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Leaves | basal sometimes 2-ranked; cauline 0–2(–3); primary leaves ternate or palmate, (0.5–)1–12(–20) cm; petiole: long hairs absent or spreading to ascending, weak to ± stiff, glands usually absent or sparse, sometimes abundant; leaflets 3–7, at tip of leaf axis, overlapping or not, broadly oblanceolate or obtriangular to flabellate, margins flat to revolute, distal 1/4–3/4+ evenly to unevenly incised 1/4–1/2 to midvein, sometimes 3–5-lobed 1/2–3/4 to midvein as well, primary teeth (1–)2–5(–7) per side or per lobe, surfaces ± similar, abaxial pale to dark green, rarely grayish, brownish, or reddish, cottony hairs absent, adaxial green to dark green, rarely brownish or reddish, sometimes ± glaucous (P. grayi), long hairs (if present) soft to weak, sometimes stiff. |
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Basal leaves | not in ranks, ternate, (1–)2–5(–7) cm; stipules: apex acute to rounded; petiole 1–4 cm, hairs absent or long hairs sparse to common, appressed, 0.5–1 mm, stiff, glands absent; leaflets 3, central obovate to flabellate, 1–2.5 × 0.5–2 cm, petiolule (1–)3–10 mm, margins flat, not lobed, distal 1/2–3/4 ± evenly incised 1/4–1/2 to midvein, teeth (2–)3(–4) per side, not secondarily toothed, surfaces similar, green, sometimes ±glaucous, hairs absent or sparse to common, 0.5 mm, glands absent. |
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Inflorescences | 1–6(–12)-flowered, ± cymose, ± to very open, or solitary flowers. |
1–6-flowered. |
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Pedicels | usually straight in fruit (often recurved in P. verna), (0–)0.2–3(–9) cm, proximal usually not much longer than distal (sometimes longer in fruit). |
straight, 0.5–2 cm in flower, to 5 cm in fruit. |
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Flowers | 5-merous; hypanthium 2–7 mm diam.; petals pale to dark yellow, mostly ± obcordate (to orbiculate in P. elegans), (2–)3–10(–12) mm, usually longer than sepals, apex usually ± retuse; stamens ca. 20; styles subapical, columnar, filiform, or tapered, not, scarcely, or ± papillate-swollen in proximal 1/5–1/2, 0.8–2.5 mm. |
epicalyx bractlets elliptic to oval, 2–4.5 × 1–2 mm, margins flat; hypanthium 2.5–4 mm diam.; sepals 3–5.5 mm, apex acute to obtuse; petals yellow, 4–7 × 4–6 mm; filaments 1–3 mm, anthers 0.5–0.9 mm; carpels 10–30, styles filiform, not or slightly papillate-swollen proximally, 1–2.5 mm. |
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Achenes | ± smooth. |
1.2–1.5 mm. |
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Potentilla sect. Aureae |
Potentilla grayi |
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Phenology | Flowering summer. | |||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Moist stream banks, lakeshores, meadows, in conifer woodlands | |||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 2000–2800 m (6600–9200 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||
Distribution | North America; Eurasia [Reportedly introduced in s Australia] |
CA |
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Discussion | Species ca. 30 (10 in the flora). The circumscription of sect. Aureae used here largely follows that of B. C. Johnston (1985), except that Potentilla sierrae-blancae is placed in sect. Concinnae and P. rubella and P. stipularis are in sect. Chrysanthae. Other treatments (for example, P. A. Rydberg 1908d; T. Wolf 1908) split the species included here in two groups and/or recombine them with species in other sections, notably sect. Ranunculoides (Th. Wolf) Juzepczuk. Since Potentilla glaucophylla (sect. Graciles) and P. cottamii (sect. Subviscosae) are sometimes identified as members of sect. Aureae, they are included herein and key out in the third and twelfth couplets, respectively. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Potentilla grayi occurs on the western slope of the Sierra Nevada from El Dorado County to Tulare County, well within the range of P. flabellifolia. Some collections are intermediate between the two. Potentilla grayi can be distinguished by its tendency to have glabrous or strigose leaves, distinctly petiolulate leaflets with simple teeth, paler yellow petals, and a less-branched caudex. The report of Potentilla grayi in New Mexico (J. D. Garcia 1970) was based on a specimen of P. norvegica (sect. Rivales). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 188. | FNA vol. 9, p. 191. | ||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | P. unranked Aureae, P. ser., P. section Frigidae | P. flabellifolia var. grayi | ||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | (Rydberg) Juzepczuk: in V. L. Komarov et al., Fl. URSS 10: 197. (1941) | S. Watson: Proc. Amer. Acad. Arts 8: 560. (1873) | ||||||||||||||||||||||||||||||||||||||||||||||||
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