Potentilla sect. Aureae |
Potentilla glaucophylla |
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blue-leaf cinquefoil, different-leaf cinquefoil, diverse-leaf cinquefoil, mountain meadow cinquefoil, vari-leaf cinquefoil |
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Habit | Perennials, tufted to cushion-forming or matted, rarely rosetted, not stoloniferous (except sometimes P. verna); taproots not fleshy-thickened; vestiture primarily of straight hairs, not differentiated into long and short, glands absent or sparse to abundant, sometimes red (P. hyparctica). | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Glands | absent or inconspicuous, uncolored. |
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Stems | usually ascending to erect, sometimes prostrate to decumbent or spreading, not flagelliform, not rooting at nodes (except P. verna), lateral to persistent basal rosettes, 0.1–2(–3) dm, lengths 1–3(–4) times basal leaves. |
mostly ± ascending, 0.5–3(–4.5) dm. |
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Leaves | basal sometimes 2-ranked; cauline 0–2(–3); primary leaves ternate or palmate, (0.5–)1–12(–20) cm; petiole: long hairs absent or spreading to ascending, weak to ± stiff, glands usually absent or sparse, sometimes abundant; leaflets 3–7, at tip of leaf axis, overlapping or not, broadly oblanceolate or obtriangular to flabellate, margins flat to revolute, distal 1/4–3/4+ evenly to unevenly incised 1/4–1/2 to midvein, sometimes 3–5-lobed 1/2–3/4 to midvein as well, primary teeth (1–)2–5(–7) per side or per lobe, surfaces ± similar, abaxial pale to dark green, rarely grayish, brownish, or reddish, cottony hairs absent, adaxial green to dark green, rarely brownish or reddish, sometimes ± glaucous (P. grayi), long hairs (if present) soft to weak, sometimes stiff. |
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Basal leaves | often 2-ranked, palmate, sometimes subpalmate, 2–20 cm; petiole (0.5–)1–8(–14) cm, long hairs often absent, sometimes sparse to abundant, usually appressed, 1–2 mm, weak to ± stiff, short, crisped, and cottony hairs absent, glands absent or sparse; leaflets 5–6(–7), sometimes with 1–2(–4) additional, smaller, entire leaflets, on tip or to distal 1/10(–1/4) of leaf axis, separate, largest ones oblanceolate or cuneate to obovate, 1–4(–6) × 0.5–1.5(–2) cm, margins flat, distal 1/3–1/2(–2/3) ± evenly incised 1/2 or nearly to midvein, undivided medial blade 1.5–7(–10) mm wide, teeth 1–3(–5) per side, broadly lanceolate or linear to narrowly oblong, 1–5(–8) mm, surfaces similar adaxial ± less hairy, ± blue-green, usually glaucous, long hairs sparse to abundant (often restricted to veins, distal teeth, and margins), short, crisped, and cottony hairs absent, glands absent or sparse. |
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Cauline leaves | (0–)1–2(–3). |
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Inflorescences | 1–6(–12)-flowered, ± cymose, ± to very open, or solitary flowers. |
2–10(–20)-flowered. |
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Pedicels | usually straight in fruit (often recurved in P. verna), (0–)0.2–3(–9) cm, proximal usually not much longer than distal (sometimes longer in fruit). |
(0.5–)1–3(–6.5) cm. |
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Flowers | 5-merous; hypanthium 2–7 mm diam.; petals pale to dark yellow, mostly ± obcordate (to orbiculate in P. elegans), (2–)3–10(–12) mm, usually longer than sepals, apex usually ± retuse; stamens ca. 20; styles subapical, columnar, filiform, or tapered, not, scarcely, or ± papillate-swollen in proximal 1/5–1/2, 0.8–2.5 mm. |
epicalyx bractlets lanceolate to elliptic, 2–5 × 0.8–1.2(–1.6) mm, hairs sparse to common, rarely abundant, ± ascending, glands absent or inconspicuous; hypanthium 3.5–5 mm diam.; sepals (2–)2.5–4.5(–5) mm, apex acute to short acuminate; petals (4–)5–10(–12) × 4–9(–10) mm; filaments 1.5–2.5(–3) mm, anthers 0.4–0.7(–0.9) mm; carpels 25–40, styles filiform above papillate-swollen base, 1.5–2.5(–3) mm. |
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Achenes | ± smooth. |
1.2–1.6 mm. |
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Potentilla sect. Aureae |
Potentilla glaucophylla |
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Distribution | North America; Eurasia [Reportedly introduced in s Australia] |
AK; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC; NL; NT; SK; YT
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Discussion | Species ca. 30 (10 in the flora). The circumscription of sect. Aureae used here largely follows that of B. C. Johnston (1985), except that Potentilla sierrae-blancae is placed in sect. Concinnae and P. rubella and P. stipularis are in sect. Chrysanthae. Other treatments (for example, P. A. Rydberg 1908d; T. Wolf 1908) split the species included here in two groups and/or recombine them with species in other sections, notably sect. Ranunculoides (Th. Wolf) Juzepczuk. Since Potentilla glaucophylla (sect. Graciles) and P. cottamii (sect. Subviscosae) are sometimes identified as members of sect. Aureae, they are included herein and key out in the third and twelfth couplets, respectively. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Potentilla glaucophylla replaces P. diversifolia; an examination of the lectotype of the latter confirms J. Soják’s (1996) conclusion that P. diversifolia applies to a hybrid involving P. glaucophylla and P. hippiana. Historically, this species was often termed P. dissecta Pursh, a misapplied name that has been rejected (B. Ertter et al. 2008). In addition to the nomenclatural change, the circumscription of Potentilla glaucophylla is here restricted to plants with usually glaucous, blue-green, distally toothed leaflets that are usually glabrate (at least in var. glaucophylla). Plants from the Colorado Plateau and southern Rocky Mountains formerly included in this species and having dark green leaves, more leaflet teeth, and larger anthers are transferred either to P. townsendii or to a currently undescribed entity. The two-ranked leaves of Potentilla glaucophylla suggest a possible connection to P. crantzii in sect. Aureae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 188. | FNA vol. 9, p. 152. | ||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | P. unranked Aureae, P. ser., P. section Frigidae | P. diversifolia var. glaucophylla | ||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | (Rydberg) Juzepczuk: in V. L. Komarov et al., Fl. URSS 10: 197. (1941) | Lehmann: Index Seminum (Hamburg) 1836: 7. (1836) | ||||||||||||||||||||||||||||||||||||||||||||||||||||
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