Potentilla sect. Aureae |
Potentilla fragiformis |
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strawberry cinquefoil |
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Habit | Perennials, tufted to cushion-forming or matted, rarely rosetted, not stoloniferous (except sometimes P. verna); taproots not fleshy-thickened; vestiture primarily of straight hairs, not differentiated into long and short, glands absent or sparse to abundant, sometimes red (P. hyparctica). | Plants tufted to cushion-forming; caudex branches short, stout, diam. 1–1.5(–2) cm, including old leaf bases. | ||||||||||||||||||||||||||||||||||||||||||||||||
Stems | usually ascending to erect, sometimes prostrate to decumbent or spreading, not flagelliform, not rooting at nodes (except P. verna), lateral to persistent basal rosettes, 0.1–2(–3) dm, lengths 1–3(–4) times basal leaves. |
ascending to erect, 0.3–1.5(–2) dm, lengths 1–2 times basal leaves. |
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Leaves | basal sometimes 2-ranked; cauline 0–2(–3); primary leaves ternate or palmate, (0.5–)1–12(–20) cm; petiole: long hairs absent or spreading to ascending, weak to ± stiff, glands usually absent or sparse, sometimes abundant; leaflets 3–7, at tip of leaf axis, overlapping or not, broadly oblanceolate or obtriangular to flabellate, margins flat to revolute, distal 1/4–3/4+ evenly to unevenly incised 1/4–1/2 to midvein, sometimes 3–5-lobed 1/2–3/4 to midvein as well, primary teeth (1–)2–5(–7) per side or per lobe, surfaces ± similar, abaxial pale to dark green, rarely grayish, brownish, or reddish, cottony hairs absent, adaxial green to dark green, rarely brownish or reddish, sometimes ± glaucous (P. grayi), long hairs (if present) soft to weak, sometimes stiff. |
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Basal leaves | not in ranks, ternate, (1–)4–10(–15) cm; stipules: apex acute to obtuse; petiole (0.3–)2–11(–13) cm, long hairs abundant, ascending to spreading, 1–2.3 mm, soft, glands sparse to common; leaflets 3, central obovate, 0.7–2.5 × 0.5–1.6 cm, petiolule 0–1 mm, margins revolute, not lobed, distal 1/2–2/3 evenly incised ± 1/3 to midvein, teeth 2–3(–4) per side, surfaces ± similar, green (paler abaxially), hairs common, 1–1.5 mm, glands common (sparser adaxially). |
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Inflorescences | 1–6(–12)-flowered, ± cymose, ± to very open, or solitary flowers. |
1–3-flowered. |
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Pedicels | usually straight in fruit (often recurved in P. verna), (0–)0.2–3(–9) cm, proximal usually not much longer than distal (sometimes longer in fruit). |
straight, 0.5–2 cm in flower, to 3.5 cm in fruit. |
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Flowers | 5-merous; hypanthium 2–7 mm diam.; petals pale to dark yellow, mostly ± obcordate (to orbiculate in P. elegans), (2–)3–10(–12) mm, usually longer than sepals, apex usually ± retuse; stamens ca. 20; styles subapical, columnar, filiform, or tapered, not, scarcely, or ± papillate-swollen in proximal 1/5–1/2, 0.8–2.5 mm. |
epicalyx bractlets oblong, 3–7 × 1.2–2.5 mm, margins revolute; hypanthium 4–6 mm diam.; sepals 4–6(–7) mm, apex broadly acute to obtuse; petals yellow, 9–12 × 8–10 mm; filaments (2–)2.8–3.2 mm, anthers 0.5–0.8 mm; carpels 40–50, styles filiform, not or scarcely papillate-swollen proximally, 1.7–2 mm. |
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Achenes | ± smooth. |
1.3–1.5. |
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2n | = 42, 56 (Russian Far East). |
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Potentilla sect. Aureae |
Potentilla fragiformis |
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Phenology | Flowering late spring–summer. | |||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Leymus-Potentilla sand dunes, coastal gravel bars, beach ridges | |||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 0–10 m (0–0 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||
Distribution | North America; Eurasia [Reportedly introduced in s Australia] |
AK; e Asia (Russian Far East) |
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Discussion | Species ca. 30 (10 in the flora). The circumscription of sect. Aureae used here largely follows that of B. C. Johnston (1985), except that Potentilla sierrae-blancae is placed in sect. Concinnae and P. rubella and P. stipularis are in sect. Chrysanthae. Other treatments (for example, P. A. Rydberg 1908d; T. Wolf 1908) split the species included here in two groups and/or recombine them with species in other sections, notably sect. Ranunculoides (Th. Wolf) Juzepczuk. Since Potentilla glaucophylla (sect. Graciles) and P. cottamii (sect. Subviscosae) are sometimes identified as members of sect. Aureae, they are included herein and key out in the third and twelfth couplets, respectively. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Recent collections of Potentilla fragiformis in Alaska on the coast of the Bering and Chukchi seas have confirmed historical reports (P. A. Rydberg 1908d) of this Beringian species in North America. Dwarf plants from St. Lawrence Island are somewhat intermediate to P. nana but have the larger petals and longer styles of P. fragiformis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 188. | FNA vol. 9, p. 191. | ||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | P. unranked Aureae, P. ser., P. section Frigidae | |||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | (Rydberg) Juzepczuk: in V. L. Komarov et al., Fl. URSS 10: 197. (1941) | D. F. K. Schlechtendal: Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk. 7: 294. (1816) | ||||||||||||||||||||||||||||||||||||||||||||||||
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