Potentilla rivalis |
Potentilla uliginosa |
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brook cinquefoil, brook or river cinquefoil, river cinquefoil, streambank cinquefoil |
Cunningham Marsh cinquefoil |
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Habit | Plants rosetted to tufted; taproots not fleshy-thickened. | |
Stems | decumbent to erect, sometimes prostrate, (0.5–)1–4(–7) dm, hairs at base not stiff, not tubercle-based, glands absent or sparse, inconspicuous. |
probably prostrate to decumbent, or ± ascending in supporting vegetation, 2.5–5.5 dm, lengths 1–2 times basal leaves. |
Leaves | ternate, palmate, or subpalmate (with distal leaflets ± confluent), basal 3–15(–25) cm, cauline 2–7(–12) cm; petiole: basal 1–9(–16) cm, cauline 0.5–4(–8) cm, long hairs usually common to abundant, ascending to spreading, 0.5–1.5 mm, usually ± weak, ± crisped hairs common to abundant, glands absent or sparse, inconspicuous; leaflets 3–5(–7), at tip to distal 1/5 (basal) or 1/2 (cauline) of leaf axis, separate to ± overlapping, largest ones oblanceolate-elliptic to obovate, (0.5–)1–5(–6) × 0.5–2(–2.5) cm, distal 1/2–3/4 of margin evenly to unevenly incised 1/3–1/2 to midvein, sometimes cleft nearly to base, teeth 3–8 per side, surfaces moderately to abundantly hairy, glands absent or sparse. |
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Basal leaves | pinnate with distal leaflets ± confluent, 15–32 × 2–4 cm; petiole 5–10(–15) cm, straight hairs absent or sparse to common, appressed, 0.5–1.5 mm, stiff, cottony hairs absent, glands absent or sparse; primary lateral leaflets 6–10(–12) per side (irregularly paired), on distal ± 1/2 of leaf axis, loosely overlapping to proximally separate, largest ones cuneate to flabellate, 1.2–2.2 × 1–3 cm, ± whole margin unevenly incised 3/4 to nearly to midvein, ultimate segments (3–)5–10(–15), linear to narrowly oblanceolate, 5–17 × 1–2.5 mm, apical tufts less than 0.5 mm, surfaces green, not glaucous, straight hairs sparse (often glabrate adaxially), appressed, 0.5–1(–1.5) mm, stiff, cottony hairs absent, glands absent or sparse. |
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Cauline leaves | 2. |
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Inflorescences | (5–)20–100+-flowered. |
6–10-flowered, very openly cymose. |
Pedicels | 0.2–1(–2) cm. |
1–6 cm, ± recurved in fruit. |
Flowers | epicalyx bractlets narrowly elliptic to ovate, (1.5–)2–4(–6) × 0.8–1.5 mm; hypanthium (2–)3–5 mm diam.; sepals 3–5 mm, apex broadly acute to obtuse; petals pale yellow to yellow, broadly oblong-obovate, 1.5–2 × 1 mm; stamens (5–)10(–15), filaments 0.4–0.9 mm, anthers 0.2–0.3 mm; carpels 40–100, styles 0.5–0.6 mm. |
epicalyx bractlets narrowly elliptic to oblong-ovate, 4–6 × 1–2.5 mm; hypanthium 5–6 mm diam.; sepals 4–7 mm, apex acute; petals 6–10 × 5–8 mm; filaments (1.5–)2–3 mm, anthers 0.7–1.2 mm; carpels ca. 10, styles 2.5–3.5 mm. |
Achenes | yellowish, 0.7–0.9 mm, ± smooth, without a corky protuberance. |
2–2.6 mm, smooth, ± carunculate. |
2n | = 14, 70. |
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Potentilla rivalis |
Potentilla uliginosa |
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Phenology | Flowering spring–summer. | Flowering spring–summer. |
Habitat | Moist meadows, stream banks, lakeshores, gravel bars in flood plains, drying marshes, open areas in river-bottom forests | Permanent oligotrophic wetlands |
Elevation | 200–2400 m (700–7900 ft) | 30–40 m (100–100 ft) |
Distribution |
AZ; CA; CO; IA; ID; IL; KS; MA; MD; ME; MN; MO; MT; ND; NE; NM; NV; NY; OK; OR; SD; TX; UT; VA; WA; WI; WY; AB; BC; MB; ON; SK; Mexico (Baja California)
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CA |
Discussion | Potentilla rivalis is most abundant in central and western North America; reports of occurrences in more eastern states (including those listed here) need confirming, as P. rivalis and P. norvegica often have been confused. Potentilla leucocarpa Rydberg was provided as a superfluous replacement name for P. millegrana; specimens annotated by Rydberg with this name include both P. biennis and P. rivalis. Potentilla rivalis is sometimes divided into three species or varieties (for example, H. J. Scoggan 1978–1979). In a strict sense, var. rivalis has subpalmately compound (5-foliate) leaves. Variety milligrana, the most common phase, has 3-foliate leaves. Plants with both 3- and 5-foliolate leaves are var. pentandra, which also tends to have five stamens, though this latter character is not correlated with the leaf features. On the Great Plains, where var. pentandra is confined, all three expressions are often found in a single population (R. L. McGregor 1986b). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Potentilla uliginosa is known only from a handful of historical collections from Cunningham Marsh, one of three wetlands in southern Sonoma County that harbor a suite of disjunct and regionally rare species (B. C. Johnston and B. Ertter 2010). Although previously included in P. hickmanii, P. uliginosa differs in having woodier caudices, longer stems and leaves, and more deeply incised leaflets occupying less of the leaf axis. Potentilla uliginosa is presumed extinct; however, a morphologically comparable collection of the P. millefolia complex from Josephine County, Oregon (Deer Creek, near Selma, 14 June 1929, Applegate 5735, UC), opens the possibility of additional undiscovered populations in the mountains of northwestern California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 139. | FNA vol. 9, p. 175. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Rivales | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae |
Sibling taxa | ||
Synonyms | P. millegrana, P. pentandra, P. rivalis var. millegrana, P. rivalis var. pentandra | |
Name authority | Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 437. (1840) | B. C. Johnston & Ertter: J. Bot. Res. Inst. Texas 4: 14, fig. 1. (2010) |
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