Potentilla rivalis |
Potentilla rubricaulis |
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brook cinquefoil, brook or river cinquefoil, river cinquefoil, streambank cinquefoil |
red-stem cinquefoil, Rocky Mountain cinquefoil |
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Caudex branches | not sheathed with marcescent whole leaves. |
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Stems | decumbent to erect, sometimes prostrate, (0.5–)1–4(–7) dm, hairs at base not stiff, not tubercle-based, glands absent or sparse, inconspicuous. |
ascending to nearly erect, 1.5–4 dm. |
Leaves | ternate, palmate, or subpalmate (with distal leaflets ± confluent), basal 3–15(–25) cm, cauline 2–7(–12) cm; petiole: basal 1–9(–16) cm, cauline 0.5–4(–8) cm, long hairs usually common to abundant, ascending to spreading, 0.5–1.5 mm, usually ± weak, ± crisped hairs common to abundant, glands absent or sparse, inconspicuous; leaflets 3–5(–7), at tip to distal 1/5 (basal) or 1/2 (cauline) of leaf axis, separate to ± overlapping, largest ones oblanceolate-elliptic to obovate, (0.5–)1–5(–6) × 0.5–2(–2.5) cm, distal 1/2–3/4 of margin evenly to unevenly incised 1/3–1/2 to midvein, sometimes cleft nearly to base, teeth 3–8 per side, surfaces moderately to abundantly hairy, glands absent or sparse. |
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Basal leaves | often both ternate and palmate on same plant, rarely subpalmate, 4–10 cm; petiole 2.5–7 cm, long hairs sparse to common, loosely appressed to ascending-spreading, 1–2 mm, ± weak to stiff, verrucose, short and/or ± crisped hairs common to abundant, cottony hairs absent, glands usually sparse; leaflets 3–5, proximalmost separated by 0(–1) mm, central oblong to obovate, 1.5–4 × 1–2.5 cm, petiolules 0–5 mm, distal 2/3–3/4 of margin incised 1/2–3/4, rarely +, to midvein, teeth (4–)5–8 per side, 4–5 mm, apical tufts 1 mm, abaxial surfaces gray to grayish white, long hairs abundant, cottony-crisped hairs usually dense, short hairs and glands absent or obscured, adaxial green to grayish green, long hairs sparse to common, 0.5–1.5 mm, stiff, short hairs absent or sparse, rarely common, crisped and cottony hairs absent, glands absent or sparse, rarely common. |
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Cauline leaves | 2–3. |
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Inflorescences | (5–)20–100+-flowered. |
4–20-flowered, open, branch angle (10–)20–45°. |
Pedicels | 0.2–1(–2) cm. |
0.5–3 cm, proximal to 5 cm. |
Flowers | epicalyx bractlets narrowly elliptic to ovate, (1.5–)2–4(–6) × 0.8–1.5 mm; hypanthium (2–)3–5 mm diam.; sepals 3–5 mm, apex broadly acute to obtuse; petals pale yellow to yellow, broadly oblong-obovate, 1.5–2 × 1 mm; stamens (5–)10(–15), filaments 0.4–0.9 mm, anthers 0.2–0.3 mm; carpels 40–100, styles 0.5–0.6 mm. |
epicalyx bractlets linear to narrowly lanceolate, 3–4 × 0.8–1.2 mm; hypanthium 4–6 mm diam.; sepals 4–5 mm, apex subacute to acute, glands usually ± sparse, not obscured; petals pale yellow, not overlapping, 5–7 × (4–)5–6.5 mm, distinctly longer than sepals; filaments 0.5–1.5 mm, anthers 0.4 mm; carpels 30–60, styles 0.9–1.1 mm. |
Achenes | yellowish, 0.7–0.9 mm, ± smooth, without a corky protuberance. |
1.2 mm. |
2n | = 14, 70. |
= 56. |
Potentilla rivalis |
Potentilla rubricaulis |
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Phenology | Flowering spring–summer. | Flowering summer. |
Habitat | Moist meadows, stream banks, lakeshores, gravel bars in flood plains, drying marshes, open areas in river-bottom forests | Sandy lake and stream shores, open sandy forests, dry grassy slopes, sandy and loamy bluffs, rock crevices, scree |
Elevation | 200–2400 m (700–7900 ft) | 0–1600 m (0–5200 ft) |
Distribution |
AZ; CA; CO; IA; ID; IL; KS; MA; MD; ME; MN; MO; MT; ND; NE; NM; NV; NY; OK; OR; SD; TX; UT; VA; WA; WI; WY; AB; BC; MB; ON; SK; Mexico (Baja California)
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AK; AB; BC; NT; SK; YT
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Discussion | Potentilla rivalis is most abundant in central and western North America; reports of occurrences in more eastern states (including those listed here) need confirming, as P. rivalis and P. norvegica often have been confused. Potentilla leucocarpa Rydberg was provided as a superfluous replacement name for P. millegrana; specimens annotated by Rydberg with this name include both P. biennis and P. rivalis. Potentilla rivalis is sometimes divided into three species or varieties (for example, H. J. Scoggan 1978–1979). In a strict sense, var. rivalis has subpalmately compound (5-foliate) leaves. Variety milligrana, the most common phase, has 3-foliate leaves. Plants with both 3- and 5-foliolate leaves are var. pentandra, which also tends to have five stamens, though this latter character is not correlated with the leaf features. On the Great Plains, where var. pentandra is confined, all three expressions are often found in a single population (R. L. McGregor 1986b). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
As addressed by B. Ertter et al. (2013), the name Potentilla rubricaulis is here restricted to relatively large plants with open inflorescences occurring mainly in glaciated parts of subarctic northwestern Canada and Alaska. Plants from the Chugach Mountains of southern Alaska tend to be more conspicuously glandular than elsewhere. The distinction between Potentilla rubricaulis and large forms of P. arenosa with supernumerary leaflets is problematic. Although both species have somewhat similar petiole vestiture (long, straight, verrucose hairs and a layer of short, stiff, or curly hairs), the latter species tends to have more stiffly spreading petiole hairs and prominently petiolulate central leaflets. The octoploid chromosome count (P. M. Dansereau and E. Steiner 1956) from Great Bear Lake area, Northwest Territories, probably belongs to Potentilla rubricaulis in the narrow sense, since that is its type locality. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 139. | FNA vol. 9, p. 207. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Rivales | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Rubricaules |
Sibling taxa | ||
Synonyms | P. millegrana, P. pentandra, P. rivalis var. millegrana, P. rivalis var. pentandra | P. dissecta var. rubricaulis, P. nivea subsp. rubricaulis |
Name authority | Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 437. (1840) | Lehmann: Nov. Stirp. Pug. 2: 11. (1830) |
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