Potentilla rivalis
Potentilla pedersenii
brook cinquefoil, brook or river cinquefoil, river cinquefoil, streambank cinquefoil
not sheathed with marcescent whole leaves.
decumbent to erect, sometimes prostrate, (0.5–)1–4(–7) dm, hairs at base not stiff, not tubercle-based, glands absent or sparse, inconspicuous.
ascending to nearly erect, 0.4–2 dm.
ternate, palmate, or subpalmate (with distal leaflets ± confluent), basal 3–15(–25) cm, cauline 2–7(–12) cm;
petiole: basal 1–9(–16) cm, cauline 0.5–4(–8) cm, long hairs usually common to abundant, ascending to spreading, 0.5–1.5 mm, usually ± weak, ± crisped hairs common to abundant, glands absent or sparse, inconspicuous;
leaflets 3–5(–7), at tip to distal 1/5 (basal) or 1/2 (cauline) of leaf axis, separate to ± overlapping, largest ones oblanceolate-elliptic to obovate, (0.5–)1–5(–6) × 0.5–2(–2.5) cm, distal 1/2–3/4 of margin evenly to unevenly incised 1/3–1/2 to midvein, sometimes cleft nearly to base, teeth 3–8 per side, surfaces moderately to abundantly hairy, glands absent or sparse.
often both ternate and palmate or subpalmate on same plant, 2.5–4 cm;
petiole 1.5–2.5 cm, long hairs common to abundant, loosely appressed to ascending-spreading, 1–2 mm, weak to ± stiff, verrucose, crisped(/short) hairs absent or sparse to common, cottony hairs absent, glands sparse to common;
leaflets 3–5, proximalmost separated by 0–2 mm, central broadly elliptic to obovate, 1–1.5 × 0.5–0.9 cm, petiolules 1–2 mm, distal 2/3–3/4 of margin incised 1/2–3/4 to midvein, teeth (2–)3–4 per side, 4–6 mm, apical tufts ± 1 mm, abaxial surfaces grayish white to white, long hairs abundant (sometimes obscuring entire surface), cottony-crisped hairs abundant to dense, short hairs absent or obscured, glands sparse to common but usually obscured, adaxial grayish green to gray, long hairs sparse to abundant, 1–1.5(–2) mm, ± weak, short (short-crisped) hairs absent or sparse, rarely common, cottony hairs absent, glands absent or sparse, rarely common.
0–2.
(5–)20–100+-flowered.
(1–)3–7-flowered, open, branch angle 30–50°.
0.2–1(–2) cm.
1–2 cm, proximal to 4 cm.
epicalyx bractlets narrowly elliptic to ovate, (1.5–)2–4(–6) × 0.8–1.5 mm;
hypanthium (2–)3–5 mm diam.;
sepals 3–5 mm, apex broadly acute to obtuse;
petals pale yellow to yellow, broadly oblong-obovate, 1.5–2 × 1 mm;
stamens (5–)10(–15), filaments 0.4–0.9 mm, anthers 0.2–0.3 mm;
carpels 40–100, styles 0.5–0.6 mm.
epicalyx bractlets narrowly ovate to elliptic, 4–5 × 1–1.4 mm;
hypanthium 3–4 mm diam.;
sepals 4–6 mm, apex subacute to acute, glands sparse to common, usually not obscured;
petals pale yellow, usually not overlapping, 6–7 × 4–8 mm, distinctly longer than sepals;
filaments 1–2 mm, anthers ± 0.4 mm;
carpels 40–80, styles 0.8–0.9 mm.
yellowish, 0.7–0.9 mm, ± smooth, without a corky protuberance.
1.1–1.2 mm.
= 14, 70.
Potentilla rivalis
Potentilla pedersenii
Potentilla rivalis is most abundant in central and western North America; reports of occurrences in more eastern states (including those listed here) need confirming, as P. rivalis and P. norvegica often have been confused. Potentilla leucocarpa Rydberg was provided as a superfluous replacement name for P. millegrana; specimens annotated by Rydberg with this name include both P. biennis and P. rivalis.
Potentilla rivalis is sometimes divided into three species or varieties (for example, H. J. Scoggan 1978–1979). In a strict sense, var. rivalis has subpalmately compound (5-foliate) leaves. Variety milligrana, the most common phase, has 3-foliate leaves. Plants with both 3- and 5-foliolate leaves are var. pentandra, which also tends to have five stamens, though this latter character is not correlated with the leaf features. On the Great Plains, where var. pentandra is confined, all three expressions are often found in a single population (R. L. McGregor 1986b).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Potentilla pedersenii and P. uschakovii account for the majority of arctic populations previously included in a broadly defined P. rubricaulis. The diagnostic morphological characters between the two species can be variable and overlapping; they are treated separately in part because of differences in presumed parental combinations. Whereas P. pulchella is the probable sect. Pensylvanicae parent for both species, the putative sect. Niveae parent for P. pedersenii is P. arenosa subsp. arenosa; that of P. uschakovii is P. subvahliana. Reflecting this parentage, P. pedersenii is distinguished by caudex branches with no marcescent whole leaves, verrucose long hairs on petioles, and inflorescences with usually several relatively small flowers. In contrast, P. uschakovii often has marcescent whole leaves sheathing the caudex branches, smooth long hairs on petioles, and one- or few-flowered inflorescences with mostly larger flowers. These generalities aside, there is much variation within both species, such that each island or population group may have its own features; it is probable that both P. pedersenii and P. uschakovii have evolved from multiple hybridization events.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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