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beautiful cinquefoil, pretty cinquefoil, soft cinquefoil, whiteleaf cinquefoil

Angell's or Boulder Mountain cinquefoil, Boulder Mountain cinquefoil

Glands

usually conspicuous, red-tipped (palmate-leaved plants), sometimes absent or inconspicuous, colorless (subpalmate-leaved plants).

Stems

ascending to erect, (1.5–)3–6(–8) dm.

(0.4–)0.6–1.1 dm, lengths 2–3 times basal leaves.

Basal leaves

sometimes 2-ranked, palmate to subpalmate, 3–25(–35) cm;

petiole 1–20(–30) cm, long hairs sparse to abundant, appressed to spreading, 2–3 mm, weak to stiff, short hairs usually absent, crisped hairs absent or sparse, cottony hairs usually absent, glands sparse to abundant;

leaflets 5–7, at tip or on distal 1/10 of leaf axis, separate to slightly overlapping, largest ones ± oblanceolate, 1–6(–9) × 1–4 cm, margins flat, distal 2/3 to nearly whole length evenly incised 1/4–1/2 to midvein, undivided medial blade 6–15 mm wide, teeth 6–12 per side, ± broadly lanceolate, 1–5 mm, surfaces usually strongly dissimilar, abaxial grayish to white, long hairs sparse to common (mostly on veins), short hairs absent, cottony or crisped-cottony hairs abundant to dense, glands usually ± abundant (at least on veins), adaxial green, rarely grayish green, not glaucous, long hairs sparse to common, short, crisped, and cottony hairs usually absent, glands usually sparse to abundant.

subpinnate, 1 pair of leaflets separate from terminal leaflets, distal leaflets distinct, 1.5–5 cm;

petiole 0.5–3(–4) cm, straight hairs abundant, loosely appressed to ascending, 1–2 mm, stiff, cottony hairs absent, glands sparse to abundant;

leaflets 5, on distal (1/10–)1/5–1/4 of leaf axis, separate, proximal pair separated from others by 1–3 mm of leaf axis, central leaflets narrowly cuneate to oblanceolate, 0.5–1.5 × 0.3–0.6 cm, petiolules 1 mm, distal 1/4(–1/3) or less of margins incised 1/2 or less to midvein, teeth (0–)1(–2) per side, ± separate, 1 mm, surfaces ± dissimilar, abaxial grayish to white, straight hairs ± abundant, appressed, 1–1.5 mm, ± stiff, cottony hairs abundant to sparse, glands ± sparse or obscured, adaxial greenish, straight hairs abundant, ± appressed, 1–1.5(–2) mm, stiff, cottony hairs absent, glands sparse to abundant.

Cauline leaves

1–3.

Inflorescences

(4–)10–50(–60)-flowered.

solitary flowers or 2–3-flowered.

Pedicels

0.5–3(–4.5) cm.

(1–)1.5–3 cm.

Flowers

epicalyx bractlets narrowly lanceolate to narrowly elliptic, 3–5(–6) × 0.8–2 mm, hairs sparse to common, ± appressed to spreading, glands absent or sparse to abundant;

hypanthium (3–)4–6 mm diam.;

sepals 4–8(–9) mm, apex acuminate;

petals 6–9(–11) × 6–13 mm;

filaments 1–3.5 mm, anthers 0.8–1.1 mm;

carpels 20–40, styles filiform to tapering above papillate-swollen base, (1–)1.5–2 mm.

epicalyx bractlets lanceolate-elliptic to ovate, 2–4 × 0.8–1.5 mm;

hypanthium 3–5 mm diam.;

sepals 3–5(–6) mm, apex ± acute;

petals (4–)5–6.5 × 4–5.5 mm;

filaments 1.5–2.5 mm, anthers 0.7–1 mm;

carpels 10–30, styles 2–2.5 mm.

Achenes

1.1–1.5(–1.6) mm.

1.5–2 mm, smooth to faintly rugose.

2n

= 70, 71, 108.

Potentilla pulcherrima

Potentilla angelliae

Phenology Flowering summer. Flowering summer.
Habitat Dry meadows in grasslands, sagebrush, scrub oak, aspen and conifer woodlands Rocky subalpine meadows
Elevation (300–)800–3600 m ((1000–)2600–11800 ft) 3300–3400 m (10800–11200 ft)
Distribution
from FNA
AZ; CA; CO; CT; ID; MT; ND; NM; NV; SD; UT; WY; AB; BC; MB; ON; QC; SK
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[BONAP county map]
from FNA
UT
[BONAP county map]
Discussion

Potentilla pulcherrima is here retained provisionally in the broad sense, encompassing intergrades between morphological extremes. The most distinctive extreme has strictly palmate leaves and conspicuous red-tipped glands. Eglandular hairs are relatively sparse and spreading on epicalyx bractlets and abaxial leaflet veins, such that the veins tend to contrast with the cottony white abaxial surface. At the other extreme are plants with subpalmate leaves, no or inconspicuous glands, and densely strigose abaxial leaflet veins. The former is understood to be the core species, towards which the description is accordingly weighted. The latter in turn represents introgression with P. hippiana (sect. Leucophyllae) or other species. The subpalmate extreme includes the lectotype of P. pulcherrima (designated by J. Soják 1996), so if a narrower circumscription of the species is adopted then either P. filipes would become the accepted name of the core species or predominant current usage of P. pulcherrima would have to be conserved by designating a different type.

By whatever circumscription, Potentilla pulcherrima is one of the more common members of sect. Graciles in the Rocky Mountains, extending east into the northern prairies. The species grows mainly in the mountains from southeastern British Columbia and southern Alberta to northern and eastern Arizona and southern New Mexico. The native range extends at least to Manitoba and North Dakota, with outlying populations in the Black Hills of South Dakota. Populations are also scattered in the mountains of eastern Nevada and disjunctly in the White Mountains of California. Adventive occurrences in fields and disturbed sites are reported from as far east as Connecticut; see discussion of P. gracilis for reports from New Hampshire. All known specimens from Minnesota and Oregon have been reidentified as particularly hairy variants of P. gracilis in the broad sense; reports from Washington are suspect.

Potentilla pulcherrima has often been treated as a variety of P. gracilis, sometimes with the filipes extreme included in var. gracilis (B. Boivin 1967–1979). The current decision to treat P. pulcherrima as a species is in part because it has a monsoonal Rocky Mountain rather than a Pacific Northwest center of distribution.

Plants combining the vestiture of Potentilla pulcherrima, including abundant glandularity, with subpalmate leaves approaching P. hippiana (leaflets on about one-fifth of leaf axis) have been named P. gracilis var. hippianoides S. L. Welsh & N. D. Atwood.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Potentilla angelliae is known only from Boulder Mountain on the Aquarius Plateau in Garfield and Wayne counties. Comparable plants, whose status has yet to be resolved, have been collected in Montezuma County, Colorado. S. L. Welsh et al. (1993) suggested that P. angelliae might best be treated as a variety of P. concinna; the combination of subpalmate leaves, tridentate leaflets, and relatively few-flowered inflorescences extending beyond the leaves makes P. angelliae a well-defined species with no evidence of intergradation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 158. FNA vol. 9, p. 180.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Graciles Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Concinnae
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Synonyms P. filipes, P. gracilis var. filipes, P. gracilis var. pulcherrima, P. pulcherrima var. condensata, P. pulcherrima var. filipes, P. pulcherrima var. wardii
Name authority Lehmann: Nov. Stirp. Pug. 2: 10. (1830) N. H. Holmgren: Brittonia 39: 342, fig. 2. (1987)
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