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sheep cinquefoil

Beringian cinquefoil, yurtsev's cinquefoil

Habit Plants rosetted to ± matted; taproots sometimes ± fleshy-thickened. Plants ± densely tufted.
Caudex branches

stout, usually columnar, sometimes sheathed with marcescent whole leaves.

Stems

prostrate to ascending, (0.3–)0.8–2(–3.5) dm, lengths (1–)1.5–3 times basal leaves.

erect, (0.2–)0.3–1.5(–2) dm, lengths 1.5–2.5(–3.5) times basal leaves.

Basal leaves

pinnate with distal leaflets ± distinct, (1.5–)2–10(–13) × 0.7–3.5(–5) cm;

petiole 0.5–3.5(–5) cm, straight hairs sparse to abundant, sometimes absent (var. decurrens), ± appressed to ascending, 1 mm, ± stiff, cottony hairs absent, glands sparse, often obscured;

primary lateral leaflets 3–6 per side (often with additional interspersed leaflets), on distal 1/2–2/3(–3/4) of leaf axis, ± separate to ± overlapping, largest ones narrowly cuneate-oblanceolate to ± obovate, (0.3–)0.5–2(–3.5) × (0.2–)0.3–0.8(–1) cm, distal 1/4 to whole margin unevenly to pinnately (at least distal leaflets of var. ovina) incised 1/2–3/4+ to midvein, ultimate teeth 2–9(–11), linear or oblong to ovate, 1–7(–9) × 1–2 mm, apical tufts 0.5–2 mm, surfaces green to grayish, not glaucous, straight hairs sparse to abundant (sparser adaxially), sometimes absent (except on margins), loosely appressed, 0.5–2 mm, ± stiff, cottony hairs absent, crisped hairs sometimes sparse to common, glands absent or inconspicuous.

(1–)2–10(–15) cm;

petiole (0.5–)1–5(–12) cm, long hairs common to dense, ± ascending to loosely appressed, sometimes spreading, 1–2 mm, ± soft, smooth, crisped/short-cottony hairs usually sparse, sometimes absent or common, glands absent, sparse, or obscured;

leaflets separate to slightly overlapping, central obovate or obtriangular, (0.5–)1–2(–3) × (0.4–)0.8–1.5(–2) cm, sessile or subsessile, base cuneate, margins revolute, distal (1/3–)1/2–2/3(–3/4) incised 1/2–2/3(–3/4) to midvein, teeth 2–3(–4) per side, usually ± distant, surfaces somewhat to more often strongly dissimilar, abaxial yellowish or grayish white to white, long hairs 0.8–1.5 mm, cottony-crisped hairs ± dense, adaxial green to grayish green, long hairs sparse to ± abundant, other hairs usually absent.

Cauline leaves

1–2.

(0–)1(–2).

Inflorescences

(1–)2–11(–20)-flowered, usually openly cymose.

usually 1(–2)-flowered, rarely to 5-flowered.

Pedicels

(0.8–)1–2(–4) cm, straight to ± recurved in fruit.

(0.5–)2–4 cm in flower, to 5 cm in fruit.

Flowers

epicalyx bractlets linear-lanceolate to narrowly elliptic, sometimes doubled, 2–3.5(–5) × 1 mm;

hypanthium 3–5 mm diam.;

sepals 3.5–5.5(–7) mm, apex acute to obtuse;

petals 4–7(–8) × 3.5–8 mm;

filaments 1–2.5 mm, anthers 0.4–1 mm usually ± 1/2 as long as filaments;

carpels 10–20, styles 2–3 mm.

epicalyx bractlets elliptic-lanceolate to ovate, (3–)4–6(–7) × (0.8–)1.2–2(–2.5) mm, (1/2–)2/3 to as wide as sepals, margins revolute, rarely flat, red glands absent;

hypanthium 2.5–4 mm diam.;

sepals 4–6(–7) mm, apex subacute;

petals (5–)6–9 × (5–)7–9 mm, significantly longer than sepals;

filaments 1–1.3 mm, anthers 0.4–0.6 mm;

carpels 30–40, apical hairs absent, styles narrowly columnar to conic-tapered, papillate-swollen on proximal 1/5(–1/3), 0.9–1.1 mm.

Achenes

1.5–2 mm, smooth, not carunculate.

1.2–2 mm.

2n

= 28, 42, 49, 56 (Russian Far East).

Potentilla ovina

Potentilla subgorodkovii

Phenology Flowering late spring to summer.
Habitat Dry alpine heaths, exposed ridges and summits, rock outcrops, scree and talus, dry tundra, acidic and calcareous bedrock
Elevation 0–4300 m (0–14100 ft)
Distribution
from FNA
CO; ID; MT; NV; OR; UT; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CO; MT; UT; WY; AB; BC; NT; NU; YT; Asia (Russian Far East, Sakha [Yakutia])
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Potentilla ovina is here accepted as an implicit new name by J. M. Macoun for P. diversifolia var. pinnatisecta, in agreement with N. H. Holmgren (1997b). In contrast, B. C. Johnston (1980) considered the names heterotypic. Potentilla ovina has priority at the species rank over P. pinnatisecta by one month.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The name Potentilla subgorodkovii was coined for a presumed hybrid species resulting from P. crebridens × P. subvahliana; here it is applied in a collective meaning for plants combining characteristics from multiple species of the P. uniflora/villosa and P. nivea groups. Morphologically, this collective entity is much closer to the P. uniflora/villosa group than to the P. nivea group and is clearly different from primary hybrids and clones, which are often observed.

Potentilla subgorodkovii constitutes about half of what previously has been considered P. uniflora in Alaska and Yukon and is the only such entity fully confirmed south of northern British Columbia. These southern populations occur outside the range of possible parental members of the P. uniflora/villosa group and do not support the hybrid hypothesis. They may instead represent a distinct species, not yet described.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Distal leaflets: ± whole margin pinnately incised 3/4+ to midvein, teeth 4–9(–11), linear, surfaces grayish, rarely green, hairs sparse to dense.
var. ovina
1. Distal leaflets: distal 1/4–1/2(–3/4) of margin unevenly to pinnately incised 1/2–2/3(–3/4) to midvein, teeth 2–5(–7), ovate to oblong, surfaces green, hairs absent (except on margins) or sparse.
var. decurrens
Source FNA vol. 9, p. 171. FNA vol. 9, p. 204.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Niveae
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Subordinate taxa
P. ovina var. decurrens, P. ovina var. ovina
Synonyms P. diversifolia var. pinnatisecta, P. pinnatisecta, P. plattensis var. pinnatisecta
Name authority J. M. Macoun: Canad. Rec. Sci. 6: 464. (1896) Jurtzev: Bot. Zhurn. (Moscow & Leningrad) 78: 83. (1993)
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