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sheep cinquefoil

beautiful cinquefoil, pretty cinquefoil, soft cinquefoil, whiteleaf cinquefoil

Habit Plants rosetted to ± matted; taproots sometimes ± fleshy-thickened.
Glands

usually conspicuous, red-tipped (palmate-leaved plants), sometimes absent or inconspicuous, colorless (subpalmate-leaved plants).

Stems

prostrate to ascending, (0.3–)0.8–2(–3.5) dm, lengths (1–)1.5–3 times basal leaves.

ascending to erect, (1.5–)3–6(–8) dm.

Basal leaves

pinnate with distal leaflets ± distinct, (1.5–)2–10(–13) × 0.7–3.5(–5) cm;

petiole 0.5–3.5(–5) cm, straight hairs sparse to abundant, sometimes absent (var. decurrens), ± appressed to ascending, 1 mm, ± stiff, cottony hairs absent, glands sparse, often obscured;

primary lateral leaflets 3–6 per side (often with additional interspersed leaflets), on distal 1/2–2/3(–3/4) of leaf axis, ± separate to ± overlapping, largest ones narrowly cuneate-oblanceolate to ± obovate, (0.3–)0.5–2(–3.5) × (0.2–)0.3–0.8(–1) cm, distal 1/4 to whole margin unevenly to pinnately (at least distal leaflets of var. ovina) incised 1/2–3/4+ to midvein, ultimate teeth 2–9(–11), linear or oblong to ovate, 1–7(–9) × 1–2 mm, apical tufts 0.5–2 mm, surfaces green to grayish, not glaucous, straight hairs sparse to abundant (sparser adaxially), sometimes absent (except on margins), loosely appressed, 0.5–2 mm, ± stiff, cottony hairs absent, crisped hairs sometimes sparse to common, glands absent or inconspicuous.

sometimes 2-ranked, palmate to subpalmate, 3–25(–35) cm;

petiole 1–20(–30) cm, long hairs sparse to abundant, appressed to spreading, 2–3 mm, weak to stiff, short hairs usually absent, crisped hairs absent or sparse, cottony hairs usually absent, glands sparse to abundant;

leaflets 5–7, at tip or on distal 1/10 of leaf axis, separate to slightly overlapping, largest ones ± oblanceolate, 1–6(–9) × 1–4 cm, margins flat, distal 2/3 to nearly whole length evenly incised 1/4–1/2 to midvein, undivided medial blade 6–15 mm wide, teeth 6–12 per side, ± broadly lanceolate, 1–5 mm, surfaces usually strongly dissimilar, abaxial grayish to white, long hairs sparse to common (mostly on veins), short hairs absent, cottony or crisped-cottony hairs abundant to dense, glands usually ± abundant (at least on veins), adaxial green, rarely grayish green, not glaucous, long hairs sparse to common, short, crisped, and cottony hairs usually absent, glands usually sparse to abundant.

Cauline leaves

1–2.

1–3.

Inflorescences

(1–)2–11(–20)-flowered, usually openly cymose.

(4–)10–50(–60)-flowered.

Pedicels

(0.8–)1–2(–4) cm, straight to ± recurved in fruit.

0.5–3(–4.5) cm.

Flowers

epicalyx bractlets linear-lanceolate to narrowly elliptic, sometimes doubled, 2–3.5(–5) × 1 mm;

hypanthium 3–5 mm diam.;

sepals 3.5–5.5(–7) mm, apex acute to obtuse;

petals 4–7(–8) × 3.5–8 mm;

filaments 1–2.5 mm, anthers 0.4–1 mm usually ± 1/2 as long as filaments;

carpels 10–20, styles 2–3 mm.

epicalyx bractlets narrowly lanceolate to narrowly elliptic, 3–5(–6) × 0.8–2 mm, hairs sparse to common, ± appressed to spreading, glands absent or sparse to abundant;

hypanthium (3–)4–6 mm diam.;

sepals 4–8(–9) mm, apex acuminate;

petals 6–9(–11) × 6–13 mm;

filaments 1–3.5 mm, anthers 0.8–1.1 mm;

carpels 20–40, styles filiform to tapering above papillate-swollen base, (1–)1.5–2 mm.

Achenes

1.5–2 mm, smooth, not carunculate.

1.1–1.5(–1.6) mm.

2n

= 70, 71, 108.

Potentilla ovina

Potentilla pulcherrima

Phenology Flowering summer.
Habitat Dry meadows in grasslands, sagebrush, scrub oak, aspen and conifer woodlands
Elevation (300–)800–3600 m ((1000–)2600–11800 ft)
Distribution
from FNA
CO; ID; MT; NV; OR; UT; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; CT; ID; MT; ND; NM; NV; SD; UT; WY; AB; BC; MB; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Potentilla ovina is here accepted as an implicit new name by J. M. Macoun for P. diversifolia var. pinnatisecta, in agreement with N. H. Holmgren (1997b). In contrast, B. C. Johnston (1980) considered the names heterotypic. Potentilla ovina has priority at the species rank over P. pinnatisecta by one month.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Potentilla pulcherrima is here retained provisionally in the broad sense, encompassing intergrades between morphological extremes. The most distinctive extreme has strictly palmate leaves and conspicuous red-tipped glands. Eglandular hairs are relatively sparse and spreading on epicalyx bractlets and abaxial leaflet veins, such that the veins tend to contrast with the cottony white abaxial surface. At the other extreme are plants with subpalmate leaves, no or inconspicuous glands, and densely strigose abaxial leaflet veins. The former is understood to be the core species, towards which the description is accordingly weighted. The latter in turn represents introgression with P. hippiana (sect. Leucophyllae) or other species. The subpalmate extreme includes the lectotype of P. pulcherrima (designated by J. Soják 1996), so if a narrower circumscription of the species is adopted then either P. filipes would become the accepted name of the core species or predominant current usage of P. pulcherrima would have to be conserved by designating a different type.

By whatever circumscription, Potentilla pulcherrima is one of the more common members of sect. Graciles in the Rocky Mountains, extending east into the northern prairies. The species grows mainly in the mountains from southeastern British Columbia and southern Alberta to northern and eastern Arizona and southern New Mexico. The native range extends at least to Manitoba and North Dakota, with outlying populations in the Black Hills of South Dakota. Populations are also scattered in the mountains of eastern Nevada and disjunctly in the White Mountains of California. Adventive occurrences in fields and disturbed sites are reported from as far east as Connecticut; see discussion of P. gracilis for reports from New Hampshire. All known specimens from Minnesota and Oregon have been reidentified as particularly hairy variants of P. gracilis in the broad sense; reports from Washington are suspect.

Potentilla pulcherrima has often been treated as a variety of P. gracilis, sometimes with the filipes extreme included in var. gracilis (B. Boivin 1967–1979). The current decision to treat P. pulcherrima as a species is in part because it has a monsoonal Rocky Mountain rather than a Pacific Northwest center of distribution.

Plants combining the vestiture of Potentilla pulcherrima, including abundant glandularity, with subpalmate leaves approaching P. hippiana (leaflets on about one-fifth of leaf axis) have been named P. gracilis var. hippianoides S. L. Welsh & N. D. Atwood.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Distal leaflets: ± whole margin pinnately incised 3/4+ to midvein, teeth 4–9(–11), linear, surfaces grayish, rarely green, hairs sparse to dense.
var. ovina
1. Distal leaflets: distal 1/4–1/2(–3/4) of margin unevenly to pinnately incised 1/2–2/3(–3/4) to midvein, teeth 2–5(–7), ovate to oblong, surfaces green, hairs absent (except on margins) or sparse.
var. decurrens
Source FNA vol. 9, p. 171. FNA vol. 9, p. 158.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Graciles
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Subordinate taxa
P. ovina var. decurrens, P. ovina var. ovina
Synonyms P. diversifolia var. pinnatisecta, P. pinnatisecta, P. plattensis var. pinnatisecta P. filipes, P. gracilis var. filipes, P. gracilis var. pulcherrima, P. pulcherrima var. condensata, P. pulcherrima var. filipes, P. pulcherrima var. wardii
Name authority J. M. Macoun: Canad. Rec. Sci. 6: 464. (1896) Lehmann: Nov. Stirp. Pug. 2: 10. (1830)
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