Potentilla ovina
Potentilla litoralis
sheep cinquefoil
coast cinquefoil, Pennsylvania cinquefoil, potentille du littoral
prostrate to ascending, (0.3–)0.8–2(–3.5) dm, lengths (1–)1.5–3 times basal leaves.
decumbent to nearly erect, (0.4–)1–4.5(–6) dm.
pinnate with distal leaflets ± distinct, (1.5–)2–10(–13) × 0.7–3.5(–5) cm;
petiole 0.5–3.5(–5) cm, straight hairs sparse to abundant, sometimes absent (var. decurrens), ± appressed to ascending, 1 mm, ± stiff, cottony hairs absent, glands sparse, often obscured;
primary lateral leaflets 3–6 per side (often with additional interspersed leaflets), on distal 1/2–2/3(–3/4) of leaf axis, ± separate to ± overlapping, largest ones narrowly cuneate-oblanceolate to ± obovate, (0.3–)0.5–2(–3.5) × (0.2–)0.3–0.8(–1) cm, distal 1/4 to whole margin unevenly to pinnately (at least distal leaflets of var. ovina) incised 1/2–3/4+ to midvein, ultimate teeth 2–9(–11), linear or oblong to ovate, 1–7(–9) × 1–2 mm, apical tufts 0.5–2 mm, surfaces green to grayish, not glaucous, straight hairs sparse to abundant (sparser adaxially), sometimes absent (except on margins), loosely appressed, 0.5–2 mm, ± stiff, cottony hairs absent, crisped hairs sometimes sparse to common, glands absent or inconspicuous.
subpinnate to subpalmate, (3–)4–15(–25) cm;
petiole (1–)2–8(–15) cm, long hairs sparse to abundant, appressed to rarely ascending, ± 1 mm, ± stiff, short hairs absent or sparse, cottony or crisped hairs absent, glands ± sparse;
leaflets 2–3 per side, on distal 1/6–1/3(–1/2) of leaf axis, ± overlapping, terminal ones narrowly lanceolate to narrowly obovate, (1.5–)2–5(–7) × 1–2.5 cm, margins revolute, incised 3/4+ to midvein, undivided medial blade 2–10 mm wide, teeth (2–)5–8 per side, ± linear-elliptic, surfaces usually ± dissimilar, abaxial green to grayish, long hairs ± abundant especially on veins, 0.5–1.5 mm, weak, short hairs absent or sparse, cottony hairs absent, crisped hairs absent or sparse to abundant, glands ± abundant, adaxial ± green, long hairs sparse to common, ± appressed, 0.5–1.5 mm, short hairs absent or sparse, cottony hairs absent, crisped hairs absent or sparse, glands ± abundant.
1–2.
(1–)2–4.
(1–)2–11(–20)-flowered, usually openly cymose.
(1–)5–30-flowered, congested, scarcely lengthening in fruit.
(0.8–)1–2(–4) cm, straight to ± recurved in fruit.
0.2–1 cm (proximal to 3 cm).
epicalyx bractlets linear-lanceolate to narrowly elliptic, sometimes doubled, 2–3.5(–5) × 1 mm;
hypanthium 3–5 mm diam.;
sepals 3.5–5.5(–7) mm, apex acute to obtuse;
petals 4–7(–8) × 3.5–8 mm;
filaments 1–2.5 mm, anthers 0.4–1 mm usually ± 1/2 as long as filaments;
carpels 10–20, styles 2–3 mm.
epicalyx bractlets narrowly, rarely broadly, elliptic to linear, 2.5–5(–6) × 1(–2) mm, lengths ± 2/3 to equal to sepals, margins ± revolute (at least in fruit);
hypanthium 3–5 mm diam.;
sepals 3–6 mm, apex ± acute, abaxial surfaces: venation moderate, glands abundant, evident at least distally;
petals yellow, 3–5 × 2–4 mm, lengths ± equal to sepals;
filaments (0.5–)1–1.5 mm, anthers 0.3–0.5 mm;
carpels 50–100, styles papillate-swollen in proximal 1/2–3/4+, 1–1.5 mm.
1.5–2 mm, smooth, not carunculate.
± 1 mm, smooth to faintly rugose.
= 28, 56.
Potentilla ovina
Potentilla litoralis
Varieties 2 (2 in the flora).
Potentilla ovina is here accepted as an implicit new name by J. M. Macoun for P. diversifolia var. pinnatisecta, in agreement with N. H. Holmgren (1997b). In contrast, B. C. Johnston (1980) considered the names heterotypic. Potentilla ovina has priority at the species rank over P. pinnatisecta by one month.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Revolute epicalyx bractlets and visible glands on the sepal tips are the most reliable characteristics to distinguish Potentilla litoralis from P. bipinnatifida where their ranges overlap in the northern Great Plains. The former is also more likely to be decumbent-ascending and tends to darken more in drying. Intermediates occur, as well as possible hybrids with P. bimundorum, P. hippiana, and P. pensylvanica.
Rydberg described Potentilla litoralis to accommodate the coastal phase of this species, concentrated around the Gulf of St. Lawrence and along the north Atlantic Coast as far south as New Hampshire. Potentilla pectinata Rafinesque was briefly adopted as the older name (for example, Frère Marie-Victorin 1935); it is both superfluous and a later homonym (B. Boivin 1952). The homotypic synonyms P. pensylvanica var. glabrata (Lehmann) S. Watson, P. bipinnatifida var. glabrata (Lehmann) Kohli & Packer, and P. glabrella Rydberg have been misapplied to P. litoralis; the type is a specimen of P. ovina var. ovina (sect. Multijugae) (J. Soják 1996). The type of P. dissecta Pursh, traditionally applied to P. glaucophylla (sect. Graciles), is closest to this species but has palmate leaves; ambiguity has led to the name being rejected (B. Ertter et al. 2008).
The current circumscription of Potentilla litoralis includes prairie populations extending through the northern Great Plains and boreal regions into eastern Alaska; those plants have generally been called P. virgulata when not combined with P. bipinnatifida, P. multifida in the broad sense, or P. pensylvanica. Outlying populations occur at least as far south as Wyoming (including the type of P. virgulata), central Nevada (possibly as introductions), and, possibly, Colorado.
Other than the tendency of prairie plants to be somewhat larger and more erect than the coastal plants, no significant morphologic distinctions have been found to match the differences in habitat and distribution. Based on their distribution map, B. L. Kohli and J. G. Packer (1976) considered the coastal and prairie populations, as well as what is here separated as Potentilla jepsonii, to comprise a single taxon, which they called P. bipinnatifida var. glabrata (Lehmann) Kohli & Packer (misapplied).
If Potentilla litoralis is treated as a variety of another species, var. virgulata has priority at varietal rank.
Unequivocal Potentilla litoralis barely enters Alaska; the majority of collections from central, western, and northern Alaska previously placed in P. litoralis represent an undescribed taxon whose dimensions are not included in the preceding description. Significant differences include more deeply incised leaflets with fewer teeth, flat, linear epicalyx bractlets, and larger petals. Some collections from around Kotzebue with relatively wide sepals and sparse glands may indicate hybridization with P. bimundorum.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Distal leaflets: ± whole margin pinnately incised 3/4+ to midvein, teeth 4–9(–11), linear, surfaces grayish, rarely green, hairs sparse to dense. | var. ovina |
1. Distal leaflets: distal 1/4–1/2(–3/4) of margin unevenly to pinnately incised 1/2–2/3(–3/4) to midvein, teeth 2–5(–7), ovate to oblong, surfaces green, hairs absent (except on margins) or sparse. | var. decurrens |
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