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sheep cinquefoil

Drummond's cinquefoil

Habit Plants rosetted to ± matted; taproots sometimes ± fleshy-thickened.
Glands

usually absent or inconspicuous, rarely conspicuous, uncolored.

Stems

prostrate to ascending, (0.3–)0.8–2(–3.5) dm, lengths (1–)1.5–3 times basal leaves.

decumbent to nearly erect, 1.5–4.5(–6) dm.

Basal leaves

pinnate with distal leaflets ± distinct, (1.5–)2–10(–13) × 0.7–3.5(–5) cm;

petiole 0.5–3.5(–5) cm, straight hairs sparse to abundant, sometimes absent (var. decurrens), ± appressed to ascending, 1 mm, ± stiff, cottony hairs absent, glands sparse, often obscured;

primary lateral leaflets 3–6 per side (often with additional interspersed leaflets), on distal 1/2–2/3(–3/4) of leaf axis, ± separate to ± overlapping, largest ones narrowly cuneate-oblanceolate to ± obovate, (0.3–)0.5–2(–3.5) × (0.2–)0.3–0.8(–1) cm, distal 1/4 to whole margin unevenly to pinnately (at least distal leaflets of var. ovina) incised 1/2–3/4+ to midvein, ultimate teeth 2–9(–11), linear or oblong to ovate, 1–7(–9) × 1–2 mm, apical tufts 0.5–2 mm, surfaces green to grayish, not glaucous, straight hairs sparse to abundant (sparser adaxially), sometimes absent (except on margins), loosely appressed, 0.5–2 mm, ± stiff, cottony hairs absent, crisped hairs sometimes sparse to common, glands absent or inconspicuous.

sometimes 2-ranked, subpinnate to pinnate (proximal leaflets often doubled, distal leaflets confluent and/or decurrent), (4–)10–25 cm;

petiole 1–10(–15) cm, long hairs absent or sparse to abundant, appressed, 1–1.5 mm, usually stiff, short, crisped, and cottony hairs usually absent, glands absent or (regionally) abundant;

leaflets 5–9, on distal 1/10–1/2(–3/4) of leaf axis, separate to ± overlapping, largest ones obovate to cuneate, (1–)2–5 × (0.7–)1–3(–3.5) cm, margins flat, distal 1/2–3/4 unevenly, sometimes evenly, incised ± 1/2 to midvein (often with additional incisions nearly to midvein), undivided medial blade 3–15 mm wide, teeth 3–7 per side (sometimes secondarily toothed), linear to lanceolate, 3–10 mm, surfaces similar, green, not glaucous, long hairs nearly absent or sparse to common (sometimes restricted to abaxial veins), short, crisped, and cottony hairs usually absent, glands usually absent, sometimes regionally abundant.

Cauline leaves

1–2.

1–3.

Inflorescences

(1–)2–11(–20)-flowered, usually openly cymose.

3–15-flowered.

Pedicels

(0.8–)1–2(–4) cm, straight to ± recurved in fruit.

1–3(–6.5) cm.

Flowers

epicalyx bractlets linear-lanceolate to narrowly elliptic, sometimes doubled, 2–3.5(–5) × 1 mm;

hypanthium 3–5 mm diam.;

sepals 3.5–5.5(–7) mm, apex acute to obtuse;

petals 4–7(–8) × 3.5–8 mm;

filaments 1–2.5 mm, anthers 0.4–1 mm usually ± 1/2 as long as filaments;

carpels 10–20, styles 2–3 mm.

epicalyx bractlets lanceolate to narrowly elliptic, 3–5 × 1–1.6 mm, hairs usually sparse, appressed to ascending, glands usually absent, sometimes regionally sparse to common;

hypanthium (3–)4–6(–7) mm diam.;

sepals 5–10 mm, apex acute to acuminate;

petals (6–)7–12 × 5–10 mm;

filaments 2–3.5 mm, anthers 0.7–1 mm;

carpels (20–)25–40, styles filiform above papillate-swollen base, 2–3 mm.

Achenes

1.5–2 mm, smooth, not carunculate.

1.5–2 mm.

2n

= 64, 92, 96, 98–108.

Potentilla ovina

Potentilla drummondii

Phenology Flowering summer.
Habitat Moist to dry meadows and adjacent slopes, in conifer woodlands, alpine tundra communities
Elevation 300–3000 m (1000–9800 ft)
Distribution
from FNA
CO; ID; MT; NV; OR; UT; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CA; MT; OR; WA; AB; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Potentilla ovina is here accepted as an implicit new name by J. M. Macoun for P. diversifolia var. pinnatisecta, in agreement with N. H. Holmgren (1997b). In contrast, B. C. Johnston (1980) considered the names heterotypic. Potentilla ovina has priority at the species rank over P. pinnatisecta by one month.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The present circumscription of Potentilla drummondii differs from that of B. Ertter (1992, 1993), which encompassed P. breweri and P. bruceae as varieties (see 28. P. bruceae discussion for details). The description above focuses on large, eglandular, subpinnate plants that occur in mountains from the central Sierra Nevada and northern Coast Ranges of California to the Olympic Peninsula of Washington, east to southern Alberta. At least some collections previously identified as P. glaucophylla (as P. diversifolia Lehmann) from the Kenai Peninsula in Alaska can be readily accommodated within this concept, as can collections from the Cabinet Mountains in Montana. The distinction between P. drummondii and subpalmate P. glaucophylla can be vague; in general, the proximalmost leaflets of P. glaucophylla are relatively small and often entire; in P. drummondii proximalmost leaflets are similar in size and dissection to other leaflets.

Pinnate-leaved regional variants merit further attention; in particular the smaller form described as Potentilla cascadensis, and glandular plants in the northern Coast Ranges of California. Relatively small plants forming uniform populations in the southern Sierra Nevada may represent stabilized hybrids with P. breweri.

The inclusion of Potentilla bruceae and P. drummondii in sect. Graciles differs from that of J. Clausen et al. (1940) and B. C. Johnston (1980, 1985), who allied both species with P. breweri in sect. Multijugae. Although subpalmate to subpinnate leaves are anomalous in primarily palmate sect. Graciles, doubled proximal leaflets and an indistinct transition between the rachis and axis of the pinnatisect terminal leaflet in these two species suggest that this state is derived from a palmate progenitor. Habit and petioles also are more compatible with sect. Graciles than sect. Multijugae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Distal leaflets: ± whole margin pinnately incised 3/4+ to midvein, teeth 4–9(–11), linear, surfaces grayish, rarely green, hairs sparse to dense.
var. ovina
1. Distal leaflets: distal 1/4–1/2(–3/4) of margin unevenly to pinnately incised 1/2–2/3(–3/4) to midvein, teeth 2–5(–7), ovate to oblong, surfaces green, hairs absent (except on margins) or sparse.
var. decurrens
Source FNA vol. 9, p. 171. FNA vol. 9, p. 159.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Graciles
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Subordinate taxa
P. ovina var. decurrens, P. ovina var. ovina
Synonyms P. diversifolia var. pinnatisecta, P. pinnatisecta, P. plattensis var. pinnatisecta P. cascadensis, P. drummondii var. cascadensis
Name authority J. M. Macoun: Canad. Rec. Sci. 6: 464. (1896) Lehmann: Nov. Stirp. Pug. 2: 9. (1830)
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