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sheep cinquefoil

brownish cinquefoil, slender brown cinquefoil, slender cinquefoil

Habit Plants rosetted to ± matted; taproots sometimes ± fleshy-thickened.
Glands

conspicuous, usually golden.

Stems

prostrate to ascending, (0.3–)0.8–2(–3.5) dm, lengths (1–)1.5–3 times basal leaves.

ascending to nearly erect, (1.5–)3–7 dm.

Basal leaves

pinnate with distal leaflets ± distinct, (1.5–)2–10(–13) × 0.7–3.5(–5) cm;

petiole 0.5–3.5(–5) cm, straight hairs sparse to abundant, sometimes absent (var. decurrens), ± appressed to ascending, 1 mm, ± stiff, cottony hairs absent, glands sparse, often obscured;

primary lateral leaflets 3–6 per side (often with additional interspersed leaflets), on distal 1/2–2/3(–3/4) of leaf axis, ± separate to ± overlapping, largest ones narrowly cuneate-oblanceolate to ± obovate, (0.3–)0.5–2(–3.5) × (0.2–)0.3–0.8(–1) cm, distal 1/4 to whole margin unevenly to pinnately (at least distal leaflets of var. ovina) incised 1/2–3/4+ to midvein, ultimate teeth 2–9(–11), linear or oblong to ovate, 1–7(–9) × 1–2 mm, apical tufts 0.5–2 mm, surfaces green to grayish, not glaucous, straight hairs sparse to abundant (sparser adaxially), sometimes absent (except on margins), loosely appressed, 0.5–2 mm, ± stiff, cottony hairs absent, crisped hairs sometimes sparse to common, glands absent or inconspicuous.

sometimes 2-ranked, palmate, (5–)10–30 cm;

petiole (1.5–)5–25(–50) cm, long hairs sparse to abundant, appressed to spreading, 1–2 mm, ± stiff, short, crisped, and cottony hairs absent, glands ± abundant;

leaflets (5–)7–9, at tip of leaf axis, separate to slightly overlapping, largest ones oblanceolate-elliptic to narrowly obovate, (2–)5–10 × 1–4 cm, margins flat to ± revolute, distal 3/4 to nearly whole length ± evenly incised (1/4–)1/2–3/4+ to midvein, undivided medial blade 2–15(–20) mm wide, teeth (5–)7–10 per side, linear to broadly lanceolate, 3–15 mm, surfaces ± similar, abaxial paler, green, not glaucous, long hairs sparse to common, short-crisped hairs absent or sparse, cottony hairs absent, glands ± abundant.

Cauline leaves

1–2.

1–3(–4).

Inflorescences

(1–)2–11(–20)-flowered, usually openly cymose.

12–30(–70)-flowered.

Pedicels

(0.8–)1–2(–4) cm, straight to ± recurved in fruit.

0.2–1(–4) cm.

Flowers

epicalyx bractlets linear-lanceolate to narrowly elliptic, sometimes doubled, 2–3.5(–5) × 1 mm;

hypanthium 3–5 mm diam.;

sepals 3.5–5.5(–7) mm, apex acute to obtuse;

petals 4–7(–8) × 3.5–8 mm;

filaments 1–2.5 mm, anthers 0.4–1 mm usually ± 1/2 as long as filaments;

carpels 10–20, styles 2–3 mm.

epicalyx bractlets lanceolate to elliptic, 3–6 × 1–2 mm, hairs sparse to common, loosely appressed to spreading, glands common to abundant;

hypanthium 4–6(–8) mm diam.;

sepals 4–8(–9) mm, apex acute to acuminate;

petals 6–9 × 6–10 mm;

filaments 1–2 mm, anthers 0.6–1 mm;

carpels 30–60, styles filiform-tapered, papillate-swollen proximally, 1.5–2 mm.

Achenes

1.5–2 mm, smooth, not carunculate.

1.5 mm.

Potentilla ovina

Potentilla brunnescens

Phenology Flowering summer.
Habitat Vernally moist rocky flats, sagebrush benchlands and slopes, usually basaltic
Elevation 400–3100 m (1300–10200 ft)
Distribution
from FNA
CO; ID; MT; NV; OR; UT; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CO; ID; MT; OR; UT; WA; WY
Discussion

Varieties 2 (2 in the flora).

Potentilla ovina is here accepted as an implicit new name by J. M. Macoun for P. diversifolia var. pinnatisecta, in agreement with N. H. Holmgren (1997b). In contrast, B. C. Johnston (1980) considered the names heterotypic. Potentilla ovina has priority at the species rank over P. pinnatisecta by one month.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The conspicuous golden glands, dark green color, and relatively broad leaflets make Potentilla brunnescens one of the better-defined species of sect. Graciles, though it has often been treated as a variety of P. gracilis. The glands, which are often on septate stalks, are particularly conspicuous on the epicalyx bractlets, hypanthium, and sepals, and at the junction of petiole and leaflets. The species occurs from the Blue Mountains of Oregon and Washington to western Montana, northern Utah, and north-central Colorado; collections from Alaska are probably not native.

As presented here, Potentilla brunnescens encompasses two morphological extremes that might merit varietal status. Leaflets of the more common and widespread extreme are incised about halfway to the midvein; in contrast, some populations in western Wyoming and adjacent Idaho have leaflets incised 3/4 or more to the midvein. The latter have often been assigned to P. gracilis var. elmeri while reserving P. gracilis var. brunnescens for the former; this is untenable because the type of P. brunnescens is the latter form.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Distal leaflets: ± whole margin pinnately incised 3/4+ to midvein, teeth 4–9(–11), linear, surfaces grayish, rarely green, hairs sparse to dense.
var. ovina
1. Distal leaflets: distal 1/4–1/2(–3/4) of margin unevenly to pinnately incised 1/2–2/3(–3/4) to midvein, teeth 2–5(–7), ovate to oblong, surfaces green, hairs absent (except on margins) or sparse.
var. decurrens
Source FNA vol. 9, p. 171. FNA vol. 9, p. 157.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Graciles
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Subordinate taxa
P. ovina var. decurrens, P. ovina var. ovina
Synonyms P. diversifolia var. pinnatisecta, P. pinnatisecta, P. plattensis var. pinnatisecta P. gracilis var. brunnescens
Name authority J. M. Macoun: Canad. Rec. Sci. 6: 464. (1896) Rydberg: Bull. Torrey Bot. Club 28: 173. (1901)
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