Potentilla norvegica
Potentilla glaucophylla
Norwegian cinquefoil, Norwegian or rough cinquefoil, potentille de norvège, rough cinquefoil
blue-leaf cinquefoil, different-leaf cinquefoil, diverse-leaf cinquefoil, mountain meadow cinquefoil, vari-leaf cinquefoil
absent or inconspicuous, uncolored.
ascending to erect, (0.5–)2–5(–9) dm, hairs at base ± stiff, tubercle-based, glands absent or sparse, inconspicuous.
mostly ± ascending, 0.5–3(–4.5) dm.
ternate, rarely palmate, 3–15(–20) cm;
petiole 1–6(–10) cm, long hairs sparse to abundant, spreading to ascending, 1–2.5(–3) mm, usually ± stiff, ± crisped hairs absent or sparse to common, glands absent or sparse, inconspicuous;
leaflets 3(–5), at tip of leaf axis, separate to ± overlapping, largest ones broadly oblanceolate or elliptic to obovate, 1–6(–10) × 0.7–4(–5) cm, distal (1/2–)2/3–3/4+ of margin usually ± evenly incised 1/4–1/3 to midvein, teeth (3–)4–8(–15) per side, surfaces sparsely to moderately hairy, sometimes glabrate or abundantly hairy, glands mostly absent.
often 2-ranked, palmate, sometimes subpalmate, 2–20 cm;
petiole (0.5–)1–8(–14) cm, long hairs often absent, sometimes sparse to abundant, usually appressed, 1–2 mm, weak to ± stiff, short, crisped, and cottony hairs absent, glands absent or sparse;
leaflets 5–6(–7), sometimes with 1–2(–4) additional, smaller, entire leaflets, on tip or to distal 1/10(–1/4) of leaf axis, separate, largest ones oblanceolate or cuneate to obovate, 1–4(–6) × 0.5–1.5(–2) cm, margins flat, distal 1/3–1/2(–2/3) ± evenly incised 1/2 or nearly to midvein, undivided medial blade 1.5–7(–10) mm wide, teeth 1–3(–5) per side, broadly lanceolate or linear to narrowly oblong, 1–5(–8) mm, surfaces similar adaxial ± less hairy, ± blue-green, usually glaucous, long hairs sparse to abundant (often restricted to veins, distal teeth, and margins), short, crisped, and cottony hairs absent, glands absent or sparse.
(0–)1–2(–3).
5–40+-flowered.
2–10(–20)-flowered.
(0.2–)0.5–2.5(–3) cm.
(0.5–)1–3(–6.5) cm.
epicalyx bractlets ± elliptic to narrowly ovate, (3–)4–8(–13) × 1.5–3(–5) mm;
hypanthium 4–7 mm diam.;
sepals 5–8 mm, apex acute to obtuse;
petals yellow, broadly obovate, (2–)3–5 × 2–4 mm;
stamens 15 or 20, filaments 0.7–2 mm, anthers 0.3–0.5 mm;
carpels 60–150, styles 0.7–0.8 mm.
epicalyx bractlets lanceolate to elliptic, 2–5 × 0.8–1.2(–1.6) mm, hairs sparse to common, rarely abundant, ± ascending, glands absent or inconspicuous;
hypanthium 3.5–5 mm diam.;
sepals (2–)2.5–4.5(–5) mm, apex acute to short acuminate;
petals (4–)5–10(–12) × 4–9(–10) mm;
filaments 1.5–2.5(–3) mm, anthers 0.4–0.7(–0.9) mm;
carpels 25–40, styles filiform above papillate-swollen base, 1.5–2.5(–3) mm.
tan to brown, 0.8–1.3 mm, usually strongly rugose, without a corky protuberance.
1.2–1.6 mm.
= 42, 56, 63, 70.
Potentilla norvegica
Potentilla glaucophylla
Potentilla norvegica is considered native in both North America and Eurasia, with the American race occasionally recognized as subsp. hirsuta for the stiff hairs on the stems, petioles, and pedicels of most plants in the flora area. This feature and other purported differences between the two races, in addition to being relatively subtle and inconstant, can be found in both America and Eurasia, though undoubtedly at least in part as introductions. Some populations in eastern Canada with glabrous stems have been distinguished as var. labradorica (for example, M. L. Fernald 1950), but such plants typically are intermixed with hairy individuals. The Löves (Á Löve 1954; Á Löve and D. Löve 1966) have argued that all three variants should be treated as distinct species due to chromosomal differences (hirsuta 2n = 56; labradorica 2n = 42; norvegica in the narrow sense 2n = 70) and obligate apomixis. Further research is needed to determine the taxonomic validity and rank of these expressions. If treated as species, P. flexuosa antedates P. labradorica (as noted by J. Soják 1969), while P. monspeliensis, although commonly used for the American race, is based on a European type (as summarized by A. Kurtto et al. in J. Jalas et al. 1972+, vol. 13).
An even greater challenge is determining native versus introduced ranges, especially given the likelihood that both native and Eurasian populations are widespread in North America. Achenes are produced prolifically and easily dispersed, to the extent that Potentilla norvegica is a contaminant in clover and hay fields and considered a weed at least in Canada (P. A. Werner and J. D. Soule 1976).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Varieties 2 (2 in the flora).
Potentilla glaucophylla replaces P. diversifolia; an examination of the lectotype of the latter confirms J. Soják’s (1996) conclusion that P. diversifolia applies to a hybrid involving P. glaucophylla and P. hippiana. Historically, this species was often termed P. dissecta Pursh, a misapplied name that has been rejected (B. Ertter et al. 2008).
In addition to the nomenclatural change, the circumscription of Potentilla glaucophylla is here restricted to plants with usually glaucous, blue-green, distally toothed leaflets that are usually glabrate (at least in var. glaucophylla). Plants from the Colorado Plateau and southern Rocky Mountains formerly included in this species and having dark green leaves, more leaflet teeth, and larger anthers are transferred either to P. townsendii or to a currently undescribed entity.
The two-ranked leaves of Potentilla glaucophylla suggest a possible connection to P. crantzii in sect. Aureae.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Leaflets incised 1/3–1/2 to midvein, teeth broadly lanceolate; range of species. | var. glaucophylla |
1. Leaflets incised 3/4+ to midvein, teeth linear to narrowly oblong; northern Rocky Mountains. | var. perdissecta |
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