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Newberry's cinquefoil

Bruce's cinquefoil

Glands

absent or inconspicuous, uncolored.

Stems

ascending to nearly erect, (1–)2–5(–6) dm.

Basal leaves

petiole 1–3.5 cm, long hairs ± abundant, 0.5–1.5 mm, short hairs absent or sparse, rarely common;

leaflets 0.2–1 cm, lobes oblanceolate to narrowly elliptic, (0.5–)1–2 mm wide, longs hairs ± abundant, short hairs absent or sparse, glands sparse to abundant.

not in ranks, ± subpalmate to subpinnate (proximal leaflets sometimes doubled, distal leaflets confluent and/or decurrent), (2–)5–20(–28) cm;

petiole (1–)3–15(–20) cm, long hairs abundant, appressed to ascending-spreading, 1–2.5 mm, weak often flattened and twisted, sometimes grading to crisped or cottony hairs, short hairs absent, glands absent or sparse;

leaflets 5–7, on distal 1/10–1/4 of leaf axis, ± overlapping, largest ones usually ± obovate, (1–)1.5–5(–6) × 1–3.5(–4) cm, margins flat, distal 2/3–3/4 evenly to unevenly incised ± 1/2 to midvein (often with additional incisions nearly to midvein), undivided medial blade 4–20 mm wide, teeth (2–)3–6(–8) per side (sometimes secondarily toothed), ± lanceolate, 4–11 mm, surfaces ± similar, abaxial ± lighter and hairier, gray-green to grayish, rarely green, not glaucous, long hairs ± abundant, short-crisped hairs absent or sparse to common, cottony hairs usually absent, glands absent or sparse.

Cauline leaves

1–2.

Inflorescences

5–50-flowered.

Pedicels

0.5–3(–4) cm.

Flowers

epicalyx bractlets broadly lanceolate to elliptic, 1.5–4(–5) × (0.4–)0.8–1.3 mm;

sepals 2–4(–5) mm, apex ± acute;

petals (3–)4–6 × (2–)3–5 mm;

filaments 1–2(–2.5) mm, anthers 0.4–0.6 mm;

carpels 20–50.

epicalyx bractlets broadly lanceolate to elliptic, 3–6(–7.5) × 1.5–2.5(–3) mm, hairs ± common to abundant, loosely appressed to ascending, glands absent or inconspicuous;

hypanthium 3.5–5.5 mm diam.;

sepals 5–10 mm, apex acute to acuminate;

petals 5–10 × 5–10 mm;

filaments 1–2.5 mm, anthers 0.7–1.2 mm;

carpels 15–30, styles filiform-tapered, papillate-swollen proximally, 2–3 mm.

Achenes

0.9–1.2 mm.

1.4–1.6 mm.

2n

= 63–73, 79, 87, 98, 129.

Potentilla newberryi

Potentilla bruceae

Phenology Flowering summer. Flowering summer.
Habitat Moist, sandy to clayey, more or less alkaline soil, especially where seasonally inundated near streams, ponds, and lakes Dry montane meadows and adjacent slopes, in conifer woodlands
Elevation 1300–1800 m (4300–5900 ft) 1200–3700 m (3900–12100 ft)
Distribution
from FNA
CA; NV; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Potentilla newberryi grows in valley bottoms in south-central Oregon, northeastern California, and northwestern Nevada. The only specimen supposedly collected in south-central Washington (W. N. Suksdorf 2718, WTU) was gathered in 1898.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

As redefined by B. Ertter and D. Mansfield (2007), Potentilla bruceae is characterized by irregularly subpalmate, weakly villose leaves. Such plants form relatively consistent and extensive populations in dry meadows centered near Lake Tahoe in California and Nevada, extending to the central Sierra Nevada and to the Warner Mountains in Oregon. This interpretation differs from that previously suggested by Ertter (1992), in which P. bruceae was interpreted as the catchall category for hybrids between P. breweri (sect. Multijugae) and P. drummondii. All three taxa were accordingly treated as varieties of P. drummondii. D. D. Keck (in J. Clausen et al. 1940) treated P. bruceae as a subspecies of P. drummondii; B. C. Johnston (1980) concluded that its placement as a variety of P. breweri was more justified. The three entities unquestionably intergrade (with the type of P. anomalofolia M. Peck possibly one result), but not beyond what is the norm for facultatively apomictic species of Potentilla.

Chromosome numbers reported by J. Clausen et al. (1940) require verification as this species; the irregular behavior observed in meiotic material suggests that at least some of the individuals were hybrids.

See discussion of 29. Potentilla drummondii regarding leaflet dissection and the inclusion of P. bruceae in sect. Graciles.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 137. FNA vol. 9, p. 159.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Arenicolae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Graciles
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Synonyms Ivesia gracilis P. drummondii subsp. bruceae, P. drummondii var. bruceae
Name authority A. Gray: Proc. Amer. Acad. Arts 6: 532. (1865) — not P. gracilis Douglas ex Hooker 1830 Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 342. (1908)
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