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cut-leaf cinquefoil, feather cinquefoil, feather or many-leaf or Klamath cinquefoil, many leaf cinquefoil

Townsend's cinquefoil

Habit Plants ± rosetted; taproots fleshy-thickened.
Glands

absent or inconspicuous, uncolored.

Stems

usually prostrate, sometimes ± decumbent, 0.4–2(–3) dm, lengths 1–2 times basal leaves.

ascending to nearly erect, (1–)2–5(–7.5) dm.

Basal leaves

pinnate with distal leaflets ± confluent, 2–15(–20) × 1–3 cm;

petiole 0.5–2(–3) cm, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5 mm, stiff, cottony hairs absent, glands absent or sparse;

primary lateral leaflets (3–)5–13 per side, on distal 2/3–3/4+ of leaf axis, separate to overlapping, largest ones cuneate to flabellate, 0.5–1.5(–2) × 0.5–2 cm, distal 2/3 to whole margin palmately or unevenly, rarely pinnately, incised 2/3 to completely to midvein, ultimate teeth or segments (1–)2–10, linear to broadly oblanceolate, 2–10 × (0.5–)1–2 mm, apical tufts to 1 mm, surfaces green to grayish green, not glaucous, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5(–2) mm, stiff, cottony hairs absent, glands sparse to common.

usually not in ranks, palmate, rarely subpalmate, (2–)5–20(–30) cm;

petiole (1–)2–15(–25) cm, long hairs usually absent or sparse and appressed, sometimes common and spreading, 1–1.5 mm, stiff, short, crisped, and cottony hairs absent, glands absent;

leaflets 5–7(–8), usually on tip, rarely on less than distal 1/10, of leaf axis, separate, largest ones narrowly oblanceolate-elliptic, (1.5–)2–5(–6) × 0.5–1.2(–1.5) cm, margins flat, distal (1/4–)1/2–3/4 evenly incised 1/4–1/3 to midvein, undivided medial blade 4–11 mm wide, teeth (2–)4–8(–9) per side, mostly broadly lanceolate, 1–2 mm, surfaces similar, green, not glaucous, long hairs usually absent or limited to veins and margins, short, crisped, and cottony hairs absent, glands absent.

Cauline leaves

(0–)1–2.

1–3.

Inflorescences

3–6(–10)-flowered, loosely cymose, sometimes racemiform.

(5–)10–20(–25)-flowered.

Pedicels

(0.5–)1–2(–4.5) cm, ± recurved in fruit.

0.7–3(–3.5) cm.

Flowers

epicalyx bractlets ± elliptic, 2–4(–6) × 1–2(–2.5) mm;

hypanthium 3–6 mm diam.;

sepals 4–6(–8) mm, apex acute;

petals 4–8(–10) × 3–7(–9) mm;

filaments 2–3.5 mm, anthers 0.7–1 mm;

carpels 10–30, styles (1.5–)2–3 mm.

epicalyx bractlets lanceolate to elliptic, (2–)2.5–5(–6) × 0.5–1(–1.4) mm, hairs sparse, ± appressed, glands absent;

hypanthium 3.5–5 mm diam.;

sepals 4–8(–9) mm, apex acute to long acuminate;

petals 5–10 × 5–10(–12) mm;

filaments 1–3 mm, anthers (0.5–)0.6–1 mm;

carpels 15–30, styles filiform above papillate-swollen base, (1–)1.5–2 mm.

Achenes

1.5–2 mm, smooth, often ± carunculate.

1.1–1.6 mm.

Potentilla millefolia

Potentilla townsendii

Phenology Flowering spring–summer. Flowering summer.
Habitat Vernally to permanently wet meadows, moist openings in conifer forests and sagebrush, alkaline flats Edges of meadows, dry gravelly flats and slopes, in montane to subalpine conifer woodlands, mixed grassland communities
Elevation 700–2200 m (2300–7200 ft) (2300–)2700–3700 m ((7500–)8900–12100 ft)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; Mexico (Chihuahua)
[BONAP county map]
Discussion

Potentilla millefolia occurs from central Oregon to the east side of the Sierra Nevada of California, with a disjunct occurrence on the alkaline flats of Reese River Valley, Nevada. Significant variation occurs in vestiture type, leaflet dissection, and flower size, but with minimal geographic correlation. The most distinctive variant, represented by the type of P. klamathensis, has relatively long, slender, spreading, pustule-based hairs, often intermixed with shorter hairs. This vestiture type does not appear to be correlated with any other characters or geographic distribution and may vary within a population.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Although previously applied only to collections from Chihuahua, Mexico, Potentilla townsendii is here used to accommodate most plants from Arizona and New Mexico previously placed in P. glaucophylla (as P. diversifolia Lehmann). Plants have narrow, shallowly toothed leaflets; stems, petioles, and leaflet veins that are often reddish; and larger anthers than P. glaucophylla. As now circumscribed, P. townsendii is known in the United States primarily from the White Mountains of east-central Arizona and the high mountains of northern New Mexico. Whether these plants are truly conspecific with those from Chihuahua, Mexico, remains to be confirmed. The species might also occur in Colorado and Utah, but the best disposition of these problematic populations is uncertain. Somewhat hairy plants with subpalmate leaves are interpreted as introgression from P. hippiana or other species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 174. FNA vol. 9, p. 153.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Graciles
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Synonyms P. klamathensis, P. millefolia var. klamathensis, P. plattensis var. klamathensis, P. plattensis var. millefolia
Name authority Rydberg: Bull. Torrey Bot. Club 23: 433, plate 277, figs. 1–5. (1896) Rydberg: Bull. Torrey Bot. Club 28: 174. (1901)
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